Anabarhynchus lyneborgi Ferguson & Irwin

Ferguson, David J., Irwin, Michael E. & Yeates, David K., 2013, New species of Anabarhynchus Macquart (Diptera: Therevidae) from arid and monsoon tropical Australia, Zootaxa 3680 (1), pp. 55-95 : 65-69

Anabarhynchus lyneborgi Ferguson & Irwin sp. n.

( Figures 8 , 9 , 10 )

Type material. Holotype: Male. AUSTRALIA: Western Australia: 158 km S Newman, 9 km N Kumarina Road House, malaise in wide sandy wash, 24.IV / 7.V.2003, M.E. Irwin, F.D. Parker, 638m, GPS: 24°37’08”S 117°36’08”E (MEI_173893) ( ANIC). Condition: ex - alcohol; dried using Hexa Methyl Di Silazane ( HMDS); dorsally pinned with stainless steel pin; middle right tibia missing, middle left leg missing, reasonable condition.

Paratypes: 17 Males, 49 Females. AUSTRALIA: Western Australia: 123, 38Ƥ, same data as holotype (MEI_173892, 173894-173898, 173900-173904, 173907, 173908, 173910-173946) ( ANIC); 8Ƥ, 82km south junction Karijini Dr on Great Northern Hwy, malaise in wash with pools, M.E. Irwin, F.D. Parker 694m, GPS: 23°07’03”S 119°05’05”E (MEI_173949-173956) ( ANIC); 23, 1Ƥ 30km ESE Three Rivers Station, malaise across dry creek bed of Gascoyne River., M.E. Irwin, F.D. Parker 504m, GPS: 25°13’06”S 118°56’09”E (MEI_173959, 173905, 173906) ( ANIC); 13, 2Ƥ 25km N Marble Bar, malaise across drying pool in Coongan River bed, M.E. Irwin, F.D. Parker, 136m, GPS: 20°56’02”S 119°51’00”E (MEI_173899, 173947, 173948) ( ANIC); 13 30km E Marble Bar at Yandicoogina Creek malaise near damp area in sandy wash, M.E. Irwin, F.D. Parker, 212m, GPS: 21°11’00”S 120°01’7”E (MEI_173909) ( ANIC); 13 Cape Range Natl Park, Milyering Ranger House, malaise below cliff near Ficus , 22m, M.E. Irwin, F.D. Parker, GPS: 22°01’08”S 113°55’09”E (MEI_163133) ( ANIC); 1Ƥ 133km SW Marble Bar, 17km E Wood-stock Station, 318m, malaise in damp wash, M.E. Irwin, F.D. Parker, GPS: 21°41’06”S 119°04’08”E (MEI_173957) ( ANIC); 2Ƥ Gregory National Park, Humbert Track 4.4km SSE Bullita Camp Ground malaise, white sand, M.E. Irwin, F.D. Parker, C. Lambkin, GPS: 16°08’52”S 130°26’28”E (MEI_173961, 173962) ( ANIC); 1Ƥ 300m Kennedy Range National Park, malaise in dry wooded Temple Gorge, M.E. Irwin, F.D. Parker, GPS: 24°39’07”S 115°10’04”E (MEI_173958) ( ANIC). Addition label on all MEI specimens; (1) CSIRO-Schlinger Pilbara Expedition, April 19–May 23, 2003. (2) Anabarhynchus lyneborgi Irwin. Det. M. E. Irwin 2010 .

Diagnosis. Frons raised; frontal setae in two narrow rows; area between frons dull-grey; frons width at anterior ocellus 3.0–4.0 x ocellus, female 3.3–4.1x ocellus. Scutum with a thin brown mid-line line within a brown band margined by dark grey with broad grey stripe either side, margined by pair of mid-dorsal grey stripes on matte dark-brown ground. Wing cell m3 open. Postspiracular pile present; katepisternum and prosternal furrow without pile. Forefemur 1 pd, hind femur 1 av; all femora black, fore tibia blackish-brown, middle and hind brown; all femora dorsally with appressed pale pile, ventral pile erect; short black setae predominantly on dorsal apical surface. The ventromedial plane of the gonocoxite with tuft of long pale pile.

Description. Male: Body length: 8.5mm. Wing length: 7mm. Head. Integument black. Frons raised; frons width at anterior ocellus 3.4x ocellus; ocellar tubercle flat; antennal base positioned low on frons, face and lower frons slightly protruding; parafacials and lateral sides of frons bright grey; frons with two narrow rows of black setae, area between lines of setae grey pubescence, at lower frons setae length is half the length of scape. Scape length 2.25x width; scape and pedicel grey, 1st segment of flagellum pale brown with a few short black setae to basal dorsal third, flagellar style pale brown one third flagellum. Occiput convex with bright-grey pubescence, several indistinct rows of variable strength pale macrosetae with one black, 32 to each side; postocciput to gena grey with long, pale, hair-like pile; ventral corner of eye with distinct swelling. Palpus pale brown, with long pale hair-like pile; labellum pale brown; prementum without black setae. Thorax. Integument brown. Scutal chaetotaxy black (pairs): np, 3; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with a thin brown mid-line line within a brown band margined by dark grey with broad grey stripe either side, margined by pair of mid-dorsal grey stripes; areas pre and post transverse suture with matte dark-brown; scutal surface with short appressed pale yellowish setae. Postspiracular pile present; katepisternum and prosternal furrow without pile; pleura and coxae grey pubescence, coxae with long pale pile admixed black macrosetae. Wing. Cell m3 open; hyaline with pale brown tint; veins and stigma brown; costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel buff brown; scabellum dorsally and ventrally white. Legs. Forefemur with 1 pd; hind femur with 1 av; all femora black with grey pubescence; all femora dorsally with appressed pale pile, ventral pile erect; short black setae predominantly to dorsal apical surface; fore tibia and basitarsus dark brown, hind tibia yellow-brown darkening apically, basitarsus dark. Abdomen. Integument yellow-brown and brown; laterally compressed; anterior bands dark brown covered with sparse appressed black pile; tergies 2–4 posterolaterally grey with appressed pale setae; tergites 2–3 posterior marginal bands viewed posteriorly appear bright white, viewed anteriorly appear grey; tergies 5–7 with weak black appressed pile; epandrium mottled brown with grey pubescence with pale setae to posterior margin; gonocoxite dark grey with outer gonocoxal processes yellow-brown. Terminalia. Paratype male (MEI_173904): epandrium ( Fig. 10 a), twice as wider than long, narrowing posteriorly, setae weak evenly distributed. Gonocoxite ( Fig. 10 b), semi-spherical slightly longer than wide when viewed ventrally, posterior ventral edge with broad and long lobe shaped outer gonocoxal process; ventromedial plane of the gonocoxite with tuft of long, pale pile, arising from a raised sclerite. Gonocoxite joined by hypandrium and fussed along a length of ventromedial plane. Gonocoxal apodeme short. Inner gonocoxal process, slightly ventrally directed and curving inward, lateral and inward edges flared along length with pale setae to apex. Gonostylus elongate shorter than inner gonocoxal process, setae to inner surface. Ventral lobe triangular. Aedeagus ( Fig. 10 c, d): distiphallus short, curved ventrally; sub-apical edge thickened posteriorly; apical end narrow and reflexed anteriorly; parameral sheath darkly sclerotised; dorsal apodeme of parameral sheath broadly triangular; posterior edge darkly sclerotised, with lateral edges dorsally directed. Ventral apodeme narrow, flared widely apically. Lateral ejaculatory apodeme widest dorsally narrowing ventrally, band like. Ejaculatory apodeme long, weakly connected, total length broadly flared laterally, apical end wide.

Variation. Male. Body length range: 7–8mm. Wing length range: 6–7mm. Frons width at anterior ocellar 3.0– 4.0x ocellus; occiput with 30–40 macrosetae to each side; middle tibia same as hind tibia. Female. Body length range: 7–9.5 mm. Wing length range: 6.5–8.5mm. Frons width at anterior ocellus 3.3–4.1x ocellus. Paratype female (MEI_173921): sternite 8 ( Fig. 10 f), roundish in shape; darkly sclerotised; anteriorly tapered with anterior margin slightly concave; posteriorly round with darkly sclerotized drop-shaped lobe inserted into the main sclerite with area of surrounding thinly sclerotised; short black setae evenly distributed anteriorly and laterally of thinly sclerotised area, and to drop-shaped lobe. Furca ( Fig. 10 e), posterior frame open; middle frame with thinly sclerotised lobe to lateral edge; anteroventral lobe elongate, curved posteriorly in line with frame.

Etymology. The specific epithet is derived to honor the late Dr. Leif Lyneborg taxonomic researcher of Anabarhynchus and the allied group.

Comments. From a large series of specimens collected over wide area of the Pilbara , north-west Western Australia, during the CSIRO-Schlinger Pilbara Expedition from April 19– May 23, 2003.

Keys to couplet 114 in Lyneborg (2001); readily separated from A. annulatus Lyneborg by having scutum distinctively marked with a pair of mid-dorsal bright grey strips on matte dark brown ground; and having two narrow rows of generally strong black setae to the frons.

The frons setae, laterally compressed abdomen, posterior lobe of sternite 8 deeply inserted into main sclerite and surrounded by thinly sclerotised area, and the ventromedial plain of the gonocoxite with tufts of long pale pile, places A. lyneborgi sp. n., in the ocypteraeformis species-group ( Lyneborg, 2001).

Figures

FIGURE 8. Anabarhynchus lyneborgi sp. n., holotype male, (MEI _ 173893): lateral view.

FIGURE 9. Anabarhynchus lyneborgi sp. n., holotype male, (MEI _ 173893): oblique view.

FIGURE 10. Anabarhynchus lyneborgi sp. n., paratype male terminalia (MEI _ 173904): A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view; paratype female (MEI _ 173921): E, furca; F, female sternite 8, ventral surface. Scale line = 0.5 mm.