Exetastes Gravenhorst, 1829

Reshchikov, Alexey & Alvarado, Mabel, 2022, Two new species of Exetastes (Hymenoptera: Ichneumonidae: Banchinae) from the Peruvian Andes, European Journal of Taxonomy 806 (1), pp. 52-63 : 55

Genus Exetastes Gravenhorst, 1829

Exetastes Gravenhorst, 1829: 395 . Type species: Ichneumon fornicator Fabricius, 1781 , by subsequent designation ( Westwood 1840: 60).

Leptobatus Gravenhorst, 1829: 432 . Type species: Leptobatus zeigleri Gravenhorst, 1829 , by subsequent designation ( Viereck 1914: 83).

Semnophrys Förster, 1869: 158 . Type species: Exetastes notatus Holmgren, 1860 , by subsequent designation ( Perkins 1962: 451).

Rhimphalea Förster, 1869: 202 . Type species: Rhimphalea brevicorpa Davis, 1897 (= bioculatus Cresson, 1872), by subsequent inclusion ( Davis 1897: 274).

Icyona Cameron, 1903: 340: Type species: Icyona rufipes Cameron, 1903 (= E. longipes (Smith, 1878)) , by monotypy.

Allexetastes Kokujev, 1904: 106. Type species: Exetastes (Allexetastes) komarovi Kokujev, 1904 , by subsequent designation ( Viereck 1914: 8).

Rhynchexetastes Cameron, 1906: 102. Type species Rhynchexetastes violaceipennis Cameron, 1906 , by monotypy.

Tegona Morley, 1913: 251. Type species: Tegona rufipes Morley, 1913 (= E. longipes (Smith, 1878)) , by monotypy.

Pseudexetastes Meyer, 1927: 308, Type species: Pseudexetastes diakonovi Meyer, 1927 , by original designation.

Diagnosis (updated from Gauld et al. 2002; Watanabe 2020)

Moderate to large body size wasps. Metasoma usually slightly posteriorly compressed. Clypeus in lateral view from weakly convex basally, to almost pyramidal, without a subapical median swelling, in anterior view, not strongly transverse, margin usually fairly evenly sclerotized but thin, slightly concave. Mandibles weakly and evenly tapered apically, lower tooth equal in length, or slightly shorter or slightly longer than upper tooth; both teeth pointed or with the apex of the upper tooth broad and obliquely chisel-shaped ( Townes 1970; Khalaim & Ruiz-Cancino 2012). Occipital carina dorsally complete, its lower part joining hypostomal carina a little above base of mandible. Antenna slender. Scape apically obliquely truncated. Mesosoma short, with the epicnemium sloping slightly backwards ventrally. Pronotum short, anteriorly with a transverse groove before broadly concave hind margin, its upper hind corner bluntly lobed, covering spiracular sclerite. Epomia absent or present as short trace-like ridge. Notauli absent. Mesopleuron with subalar prominence present as a low rounded to sharp promontory. Epicnemium without a distinct vertical tooth-like lamella near lower corner of pronotum. Posterior transverse carina of mesosternum entirely absent. Metanotum with hind rim simple or with a small tooth-like projection posteriorly. Metapleuron with submetapleural carina narrow, only slightly and evenly broadened anteriorly. Metasternum with weak ridges between coxal insertions. Propodeum short and quite steeply declivous.All propodeal carinae (except sometimes the pleural carina) absent. Propodeal spiracle subcircular to slightly oval. Legs slender with tibial spurs long. Hind leg very long and strong, with enlarged coxa. All tarsomeres cylindrical. Tarsal claws simple. Mid tibia without distinct denticles on outer surface. Fore wing with a large kiteshaped areolet. 2m-cu slightly sinuous, with single but long bulla. Usually cu-a opposite or slightly distal to base of Rs and M. Hind wing with distal abscissa of Cu1 joining cu-a close to M. Upper outer corner of subbasal cell almost right-angled, 95–100°. T1 with spiracle in anterior or central part. Glymma vestigial. Sclerotized part of first sternite not reaching spiracle. T2 slightly elongate (l.l–1.4 times as long as broad), with broad shallow thyridium anteriorly. Laterotergites of T2–3 indistinct, laterotergites of T4–5 not distinctly separated. Female hypopygium rather large and completely selerotized. Ovipositor short to moderately long (0.3–1.3 times as long as hind tibia), usually curved downward, compressed basally, its apex usually with a distinct dorsal subapical notch, without teeth on distal end of upper valve; rarely apically elongate tapered with a vestigial apical notch. Male with genital capsule short with a broad shallow notch ventrally.