Anthobium deplanatum Shavrin & Smetana, 2019

Shavrin, Alexey V. & Smetana, Aleš, 2019, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group, Zootaxa 4688 (4), pp. 451-482 : 474-476

Anthobium deplanatum Shavrin & Smetana , sp.n.

( Figs. 74 , 83–84 , 87 )

Type material examined: Holotype ♂: ‘ CHINA: W-Sichuan | Ya`an Prefecture, Tianquan | co., E Erlang Shan

Pass’ <rectangular label, printed>, ‘ 2900m, 20.VI.1999 | 29.52.36N, 102.17.82E | leg. A. Pütz’ <rectangular label, printed>, ‘ Sammlung | Andreas Pütz | Eisenhüttenstadt’ <yellow rectangular label, printed>, ‘ HOLOTYPE | Anthobium | deplanatum sp.n. | Shavrin A. & Smetana A. 2019’ <red rectangular label, printed> (temporarily in CAP, to be eventually deposited in SNSD).

Paratypes (2 specimens): 1 ♂ [dissected], 1 ♀: ‘ CHINA: W-Sichuan 20. VI. 1999 | Ya`an Prefecture , Tianqan Co. | E Erlang Shan Pass, 2900 m | 9 km SE Luding, 29°52N, | 102°18E, Bachufer, Moos + | Schotter, leg. M. Schülke’ <rectangular label, printed>, ‘ Sammlung | M. Schülke | Berlin’ <green rectangular label, printed> (1 ♂: CS; 1 ♀: CSC) ; 1 ♀ [Specimen without right elytron and right hind tarsus]: ‘ CHINA: W-Sichuan | Ya`an Prefecture , Tianquan | Co., E Erlang Shan Pass’ <rectangular label, printed>, ‘ 2900m, 20.VI.1999 | 29.52.36N, 102.17.82E | leg. A.Pütz’ <rectangular label, printed>, ‘ Sammlung | Andreas Pütz | Eisenhüttenstadt’ <yellow rectangular label, printed> ( CAP). All paratypes with additional red rectangular printed label : ‘ PARATYPE | Anthobium | deplanatum sp.n. | Shavrin A.V. & Smetana A. 2019’.

Description. Measurements (n=4): HW: 0.80–0.82; HL: 0.52–0.55; AL (holotype): 1.86; OL: 0.20–0.22; PL: 0.62–0.66;PW:1.35–1.40; ESL:1.52–1.74; EW:1.80–2.27;AW:1.45–1.65; MTbL(holotype):0.85,MTrL(holotype): 0.35 (MTrL 1–4: 0.18; MTrL 5: 0.17); AedL(paratype): 0.82; TL: 3.63–4.50(holotype).

Body and antennomeres 4–11 reddish-brown; mouthparts, antennomeres 1–3, lateral portions of pronotum and elytra yellow to yellow-brown. Middle portion of head with indistinct, irregular, transverse meshes between punctures, and dense, transverse microsculpture on posterior portion of infraorbital ridges; neck with isodiametric microsculpture; pronotum and scutellum without meshes. Head with fine, moderately deep punctation, irregular and sparser in middle portion between eyes, markedly coarser and denser on infraorbital ridges, with small paired impunctated portions in middle at level of posterior third of eyes; neck with irregular, fine and deep punctures; pronotum with dense, irregular punctation, finer and denser on medioapical and mediobasal portions, sparser in middle (holotype with impunctated area in mediobasal third), markedly denser, larger and deeper in lateral portions; scutellum with several fine punctures; punctation of elytra denser and finer on prescutellar area, each elytron forming six to seven very vague and tangled longitudinal rows of punctures; abdominal tergites with indistinct fine punctation. Habitus as in Fig. 74 .

Head 1.4–1.5 times as wide as long; middle portion distinctly elevated, with moderately deep semicircular impression in middle; grooves in front of ocelli deep and short, reaching posterior third of eyes; surface between punctures on infraorbital ridges irregularly elevated. Eyes moderately large. Distance between ocelli about one and half times as long as distance between ocellus and posterior margin of eye. Antenna long, reaching one fourth length of elytra when reclined; antennomeres 3–10 elongate; length × width of antennomeres (holotype): 1: 0.20 × 0.09; 2: 0.12 × 0.06; 3: 0.16 × 0.05; 4: 0.16 × 0.06; 5–7: 0.17 × 0.06; 8–9: 0.16 × 0.07; 10: 0.16 × 0.08; 11: 0.23 × 0.09.

Pronotum distinctly more than twice as long as wide, 1.6–1.7 times as wide as head; lateral sides gradually narrowed anteriad, with moderately narrowly protruded anterior angles; posteriolateral margins of pronotum distinctly emarginated in front of acute posterior angles; lateral edges of pronotum with irregular, small, rounded crenulation (mediolateral margin of pronotum of one paratype with two small indistinct protrusions); middle elevated portion with two irregular, wide and moderately deep, transverse impressions in medioapical and mediobasal portions, holotype and one paratype with additional moderately wide and deep longitudinal impression between transverse impressions; middle elevation with moderately deep, transverse impression connecting with mediolateral pits.

Elytra markedly wider than long, distinctly more than twice as long as pronotum, distinctly widened apicad from middle; surface of each elytron with indistinct three to four longitudinal elevation between punctures in middle.

Male. Protarsomeres 1–4 slightly widened. Apical margins of abdominal tergite VIII and sternite VIII emarginated. Aedeagus ( Fig. 83 ) semioval, with widely rounded apex of median lobe; parameres moderately narrow, slightly exceeding apex of median lobe, with two apical and two preapical setae; internal sac very long and complicated, with several fields of long sclerotized thorns in middle portion. Aedeagus laterally as in Fig. 84 .

Female. Protarsomeres 1–4 moderately narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded.

Comparative notes. Regarding the body size and the length of the elytra, A. deplanatum sp.n. is similar to A. kashmiricum , from which it can be distinguished by the darker coloration, shapes of the anterior angles of the pronotum, and by the external and internal structures of the aedeagus.

Distribution. The species is known only from the type locality in Erlang Shan range in Sichuan, China ( Fig. 87 ).

Bionomics. The specimens were collected at elevation 2900 m a.s.l. by sifting of mosses and gravel. Etymology. The specific epithet is the Latin adjective deplanatus, - a, - um (flattened). It refers to the markedly flattened forebody of the species.

Figures

FIGURE 87. Distribution of Anthobium deplanatum (black circles), A. gaoligongshanense (black square) and A. transversale (black triangle) in the south-western China.

FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm.

FIGURES 83–86. Aedeagus of Anthobium deplanatum (83–84: paratype) and A. gaoligongshanense (85–86: holotype): 83, 85—parameral view, 84, 86—lateral view. Scale bar: 0.1 mm.