Anthobium kashmiricum ( Cameron, 1941 )

Shavrin, Alexey V. & Smetana, Aleš, 2019, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group, Zootaxa 4688 (4), pp. 451-482 : 477-478

Anthobium kashmiricum ( Cameron, 1941)

( Figs. 76 , 80–82 )

Lathrimaeum (Prionothorax) kashmiricum Cameron, 1941: 58

Anthobium kashmiricum: Herman, 2001: 233

Anthobium (Prionothorax) kashmiricum: Smetana, 2001: 239 ; Schülke & Smetana, 2015: 307

Type material examined: Holotype by monotypy (“Type in my collection”) of Lathrimaeum kashmiricum Cameron, 1941 [specimen was dissected and reglued on a new plate; old plate under a new; aedeagus in small vial with glycerine is pinned under old plate; abdominal tergite VIII, sternite VIII and apical segment was glued on the same plate under the specimen; left hind tarsus is glued under hind leg; additional barcode label: ‘ NHMUK 013684152’] ♂: ‘SYN- | TYPE’ <round printed label with blue margin>, ‘Type’ <round printed label with red margin>, ‘Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’ <rectangular printed label>, ‘M.Cameron | Bequest. | B.M. 1955-147.’ <rectangular printed label>, ‘L. | kashmiricum | TYPE [in red] Cam.’ <rectangular label, handwritten in black Indian ink>, ‘ Anthobium | kashmiricum ( Cameron, 1941) | Shavrin A. V. 2016 ’ ( BMNH).

Additional material. INDIA: KASHMIR: 3 ♂♂, 3 ♀♀: ‘SYN- | TYPE’ <round printed label with blue margin>, Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’ <rectangular printed label>, ‘ M.Cameron | Bequest. | B.M. 1955-147.’ <rectangular printed label> ( BMNH); 2 ♂♂, 1 ♀: ‘ Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’ <handwritten in black Indian ink>, ‘ W. Steel coll. | B.M. 1969-552’ <rectangular printed label>, ‘SYN- | TYPE’ <round printed label with blue margin> ( BMNH) .

Redescription. Measurements (n=10): HW: 0.79–0.80, HL: 0.45–0.50; AL(holotype): 1.82; OL: 0.20; PL: 0.60–0.70; PW: 1.25–1.45; ESL: 1.50; EW: 1.60–1.85; AW: 1.50–1.60; MTbL(holotype) : 0.85, MTrL(holotype): 0.35 (MTrL 1–4: 0.25; MTrL 5: 0.10); AedL: 0.50; TL: 3.70(holotype)–4.45.

Body and antennomeres 4–11 (or 5–11) yellow-brown to brown, sometimes with head and bottom of elytral punctures dark-brown; lateral portions of pronotum yellow to yellow-brown; mouthparts, antennomeres 1–3 (or 1–4) and legs yellow (holotype markedly paler than other studied specimens). Head with irregular and indistinct, transverse meshes in middle of vertex, posterior portions of infraorbital ridges with rugose sculpture between punctures, middle part of neck and abdomen with isodiametric microsculpture. Head with irregular and dense punctation, markedly denser in middle portion, punctation of posterior portions of infraorbital ridges denser, with interspaces between punctures as diameter of one puncture, middle part of head behind transverse impression with narrow triangular impunctate area; pronotum with irregular, very dense and somewhat larger punctation than that on posterior part of head, more rugose in middle elevation and distinctly sparser on lateral portions; scutellum with several fine punctures; punctation of elytra as that on pronotum but sparser, middle portions of each elytron with vague and tangled six rows of longitudinal punctures, finer and more tangled in prescutellar area; abdominal tergites with regular, sparse and fine punctation. Habitus as in Fig. 76 .

Head 1.6–1.7 times as wide as long; middle part irregularly elevated, with distinct and moderately deep transverse impression, with long grooves in front of ocelli, stretching toward level of posterior third of eyes; middle part of head sometimes with indistinct transverse elevations between punctures; postocular ridges acute. Distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Antenna long, reaching apical fourth of elytra when reclined; length × width of antennomeres (holotype): 1: 0.22 × 0.10; 2: 0.15 × 0.05; 3–6: 0.17 × 0.04; 7: 0.17 × 0.06; 8: 0.15 × 0.06; 9: 0.15 × 0.07; 10: 0.13 × 0.08; 11: 0.17 × 0.08.

Pronotum 1.5–1.8 times as wide as head, widest about middle, indistinctly narrowed anteriad and slightly more narrowed posteriad; anterior angles rounded, strongly protruded anteriad; lateral margins of pronotum with irregular, indistinct crenulation; middle portion with wide longitudinal impression in middle and distinct transverse impression in mediobasal third; lateral portions moderately wide, explanate.

Elytra wider than long, markedly more than twice as long as pronotum, slightly widened posteriad from middle; lateral portions wide and explanate; surface of each elytron usually with five longitudinal elevations between punc- tures in middle, with more distinct longitudinal elevation, diagonally stretching from shoulders to posterior fourth of elytra.

Male. Protarsomeres 1–4 slightly widened. Apical margins of abdominal tergite VIII and sternite VIII straight emarginated. Aedeagus ( Fig. 80 ) small, suboval, with median lobe narrowing toward narrowly rounded apex; parameres short, slightly widened apically, hardly exceeding apex of median lobe, with two pairs of apical and preapical setae; internal sac narrow, long, spirally rolled in basal portion. Aedeagus laterally as in Fig. 81 .

Female. Protarsomeres 1–4 narrow. Apical margin of abdominal tergite VIII emarginated. Apical margin of abdominal sternite VIII widely rounded.

Comparative notes. Based on the body size and the length of the elytra, A. kashmiricum is similar to A. deplanatum sp.n., from which it can be distinguished by the paler coloration, shape of the anterior angles of the pronotum, and by the external and internal characters of the aedeagus.

Distribution. The species is known only from the type locality in Kashmir, India ( Fig. 82 ).

Bionomics. Any bionomical data are unknown.

Figures

FIGURE 82. Distribution of Anthobium reflexum (black circles) and A. kashmiricum (black squares) in Central Asia.

FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm.

FIGURES 78–81. Aedeagus of Anthobium reflexum (78–79: Kazakhstan) and A. kashmiricum (80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm.