Fusininae Wrigley, 1927

Harzhauser, Mathias, Landau, Bernard M. & Vermeij, Geerat J., 2024, The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5470 (1), pp. 1-92 : 36

Subfamily Fusininae Wrigley, 1927

Revised diagnosis. “ Members of Fusininae are characterized by elongate fusiform shells with a smooth columella, a planar to gently convex adult outer shell lip, external spiral sculpture of primary, secondary, and often tertiary elements, and axial shell sculpture consisting of rounded or rarely angular ribs, tubercles or nodes at least in early stages of postlarval growth. The teleoconch whorls are rounded to angulate in profile and are not flat-sided. The protoconch of most living species comprises one smooth whorl and a half whorl ornamented with axial riblets, but some living species of the new genus Goniofusus have a multispiral one. The aperture in fusininae is ovate to almost round; the variably elongate siphonal canal is typically straight except for a weakly dorsally deflected distal sector ” ( Vermeij & Snyder 2018: 58).

Genus Angustifusus Vermeij & Snyder, 2018

Type species. Fusus vindobonensis Hoernes & Auinger, 1890 ; original designation by Vermeij & Snyder (2018: 71). Middle Miocene , Austria .

Original diagnosis. “Shell elongate to very elongate (shell length to diameter ratio 2.7: 4.0) with small aperture and elongate, very narrowly open siphonal canal (canal length to shell length ratio 0.32: 0.42, opening less than 2 mm wide); sutures shallow and closely appressed; axial sculpture consisting of low ribs forming spirally elongate nodes at shoulder angulation, usually expressed only on spire whorls and obsolete or absent on last one or two whorls; spiral sculpture consisting of very low threads and cords, usually reduced on adult whorls; aperture adapically rounded; outer lip edge planar to abapically convex; inner side of outer lip smooth; inner lip adherent or weakly erect, usually with parietal ridge at its adapical end.” ( Vermeij & Snyder 2018: 71).

Discussion. In contrast to the original diagnosis, the outer lip of the type species may occasionally bear delicate lirae inside. Lirae are also documented in some specimens of Angustifusus hoessii ( Naumann, 1852) . This feature is variable even within a species.

Vermeij & Snyder (2018) noted that all genera within the Fusininae had paucispiral protoconchs, except Goniofusus Vermeij & Snyder, 2018 . At least Angustifusus hoessii ( Naumann, 1852) and Angustifusus aturensis ( Grateloup, 1845) have multispiral protoconchs suggestive of planktotrophic development. Again, protoconch type appears to be a variable generic character.