Drymaria villosa Schltdl. & Cham., Linnaea

Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas & Montesinos-Tubée, Daniel B., 2025, A nomenclator of Drymaria (Caryophyllaceae), PhytoKeys 266, pp. 139-175 : 139-175

54 a. Drymaria villosa Schltdl. & Cham., Linnaea 5 (2): 232. 1830. subsp. villosa

≡ Drymaria cordata var. villosa (Schltdl. & Cham.) Rohrb., Fl. Bras. 14 (2): 260. 1872.

= Drymaria hirsuta Bartl. in Presl, Reliq. Haenk. 2 (1): 8. 1831. Type: Peru: montanis huanoccensibus, 1822, T. Haenke s. n. ( lectotype, designated here: GOET [GOET 000583 image!]; isolectotype: PR [24307 image!]).

= Drymaria cubensis Regel, Allg. Gartenzeitung 8 (38): 298. 1840. Type: probably Cuba. [Probably cultivated in the Botanical Garden of Berlin from Cuban seed]. ( holotype: K [K 000471737 image!]).

= Drymaria ciliata C. A. Mey., Index Seminum (St. Petersburg) 9: 71. 1843. Type: This species was described on the basis of cultivated plants known in the horticultural trade.

= Drymaria cordata var. pilosa Schltdl., Linnaea 26: 374. 1853. Type: Mexico. This variety was described in a treatment of miscellaneous Mexican plants.

= Drymaria stylosa Backer, Bull. Jard. Bot. Buitenzorg , sér. 2 12: 15. 1913. Type: Indonesia: many localities in the mountains, s. d., C. A. Backer s. n. (without information about type specimens).

= Drymaria tepicana M. E. Jones, Contr. W. Bot. 15: 124. 1929. Type: Mexico: Nayarit: Tepic, 16 February 1927, M. E. Jones 22847 ( holotype: POM [162845 / 0002390 image!]).

≡ Drymaria villosa f. tepicana (M. E. Jones) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 227. 1961.

= Drymaria barrancae M. E. Jones, Contr. W. Bot. 18: 65. 1933. Type: Mexico. Jalisco: La barranca, Guadalajara, 17 November 1930, M. E. Jones 27051 a ( holotype POM [192857 / 0002400 image!]; isotype GH 00037718 image!]).

Type.

Mexico. Veracruz: Jalapa, without date, C. J. W. Schiede s. n. ( lectotype, designated by Athira and Maya 2025, pg. 197: LE [LE 00018305 image!]).

Distribution.

Subspecies of wide distribution throughout America and Asia.

Notes.

Drymaria villosa Cham. & Schltdl. , originally described in 1830, is equivalent to Drymaria cordata var. villosa (Cham. & Schltdl.) Rohrb. from 1872. In 1961, Duke referred to a specimen at the Komarov Herbarium (LE) as an “ isotype, ” but according to Article 9.10 of the Shenzhen Code, this should be considered a lectotype. Arya et al. (2024) accepted this correction but incorrectly designated a specimen housed at LECB as the lectotype. However, Chamisso’s original material was transferred to LE (Komarov Institute) in 1840, not to LECB, making their lectotypification invalid ( Berger 2018). Athira and Maya (2025) correctly designated specimen LE 00018305 from the Komarov Herbarium in St. Petersburg as the lectotype, as it matches Schlechtendal’s original description, ensuring nomenclatural stability. This approach follows Athira and Maya (2025) in recognizing LE 00018305 as the lectotype of D. villosa Schltdl. & Cham. Finally , although recent plastome sequencing and phylogenetic studies place D. villosa within Caryophyllaceae ( Chetri et al. 2024) , these molecular data do not affect the nomenclatural typification.