Marcusenius Gill, 1862

Kramer, Bernd & Wink, Michael, 2013, East-west differentiation in the Marcusenius macrolepidotus species complex in Southern Africa: the description of a new species for the lower Cunene River, Namibia (Teleostei: Mormyridae), Journal of Natural History (J. Nat. Hist.) 47 (35 - 36), pp. 2327-2362 : 2333-2335

Genus Marcusenius Gill, 1862

Diagnosis (translation of Taverne 1971a:106)

Body moderately elongated; snout rather low, shorter than postorbital segment of the skull and chin with mental swelling; caudal peduncle 2 to 5 times longer than deep; dorsal fin with 19 to 36 rays; anal fin with 25 to 43 rays; pectoral fin with 10 to 12 rays; 38 to 98 scales in lateral series; 8 to 26/ 12 to 28 in transversal line at the level of the body; 7 to 21/7 to 21 scales in transversal line between dorsal and anal fins; 8 to 18 scales around caudal peduncle; 3 to 8/3 to 10 conical or bicuspid teeth; lateral ethmoid present and well developed; mesethmoid small and straight; 5 circumorbital bones; pre-orbital and first infraorbital fused; 5 hypural bones; 42 to 49 vertebrae. Taverne (1971b) also gives a diagnosis including more detail on skeletal characters and a phylogeny ( Taverne 1972) that, for the purpose of the present paper, can be summarized briefly as follows: the genus Marcusenius Gill 1862 sensu stricto, that is, in Taverne’s definition that is valid at present, shares a lateral ethmoid with four other mormyrid genera, it shares five rather than six circumorbital bones with two of these other genera, but it does not share a reduced upper jaw with any of these ( Taverne 1971b; 1972).

Type species

Marcusenius cyprinoides (Linnaeus, 1758) Included species (from Eschmeyer 2013). Valid unless stated otherwise.

abadii , Gnathonemus Boulenger, 1901 . Current status: Valid as Marcusenius abadii

(Boulenger, 1901). altisambesi , Marcusenius Kramer, Skelton, Van der Bank and Wink, 2007 . angolensis , Gnathonemus Boulenger, 1905 . Current status: Valid as Marcusenius

angolensis ( Boulenger, 1905). annamariae, Gnathonemus Parenzan, 1939 . Current status: Valid as Marcusenius

annamariae (Parenzan, 1939). bentleyi , Mormyrus Boulenger, 1897 . Current status: Valid as Marcusenius bentleyi

(Boulenger, 1897). brucii , Gnathonemus Boulenger, 1910 . Current status: Valid as Marcusenius brucii

( Boulenger, 1910). cuangoanus , Gnathonemus Poll, 1967 . Current status: Valid as Marcusenius cuangoanus

(Poll, 1967). cyprinoides , Mormyrus Linnaeus, 1758 . Current status: Valid as Marcusenius cypri-

noides (Linnaeus, 1758). deboensis , Gnathonemus Daget, 1954 . Current status: Valid as Marcusenius deboensis

(Daget, 1954). devosi , Marcusenius Kramer, Skelton, Van der Bank and Wink, 2007 . dundoensis , Gnathonemus Poll, 1967 . Current status: Valid as Marcusenius dundoensis

(Poll, 1967). friteli , Gnathonemus Pellegrin, 1904 . Current status: Valid as Marcusenius friteli

(Pellegrin, 1904). furcidens , Gnathonemus Pellegrin, 1920 . Current status: Valid as Marcusenius furcidens

(Pellegrin, 1920). fuscus , Gnathonemus Pellegrin, 1901 . Current status: Valid as Marcusenius fuscus

(Pellegrin, 1901). ghesquierei , Gnathonemus Poll, 1945 . Current status: Valid as Marcusenius ghesquierei

(Poll, 1945). gracilis, Marcusenius Kramer, 2013 . greshoffii , Mormyrus Schilthuis, 1891 . Current status: Valid as Marcusenius greshoffii

(Schilthuis, 1891). intermedius, Marcusenius Pellegrin, 1924 . kainji , Marcusenius Lewis, 1974 . Uncertain status as Marcusenius kainji Lewis, 1974 .

Current status: Marcusenius kainji Lewis, 1974 . kutuensis, Gnathonemus Boulenger, 1899 . Current status: Valid as Marcusenius

kutuensis (Boulenger, 1899). leopoldianus, Gnathonemus Boulenger, 1899 . Current status: Valid as Marcusenius

leopoldianus (Boulenger, 1899). livingstonii, Gnathonemus Boulenger, 1899 . Current status: Valid as Marcusenius

livingstonii (Boulenger, 1899). multisquamatus , Marcusenius Kramer and Wink, 2013 . macrolepidotus , Mormyrus Peters, 1852 . Current status: Valid as Marcusenius

macrolepidotus ( Peters, 1852). macrophthalmus, Gnathonemus Pellegrin, 1924 . Current status: Valid as Marcusenius

macrophthalmus (Pellegrin, 1924). mento , Mormyrus Boulenger, 1890 . Current status: Valid as Marcusenius mento

(Boulenger, 1890). meronai, Marcusenius Bigorne and Paugy, 1990 .

monteiri , Mormyrus Günther, 1873 . Current status: Valid as Marcusenius monteiri (Günther, 1873) .

moorii , Mormyrus Günther, 1867 . Current status: Valid as Marcusenius moorii (Günther, 1867) .

ntemensis , Gnathonemus Pellegrin, 1927 . Current status: Valid as Marcusenius ntemensis (Pellegrin, 1927) .

pongolensis , Gnathonemus Fowler, 1934 . Current status: Valid as Marcusenius pongolensis (Fowler, 1934) .

sanagaensis, Marcusenius Boden, Teugels and Hopkins, 1997 .

schilthuisiae , Gnathonemus Boulenger, 1899 . Current status: Valid as Marcusenius schilthuisiae (Boulenger, 1899) .

senegalensis , Mormyrus Steindachner, 1870 . Current status: Valid as Marcusenius senegalensis (Steindachner, 1870) .

stanleyanus , Mormyrus Boulenger, 1897 . Current status: Valid as Marcusenius stanleyanus (Boulenger, 1897) .

thomasi , Gnathonemus Boulenger, 1916 . Current status: Valid as Marcusenius thomasi (Boulenger, 1916) .

ussheri , Mormyrus Günther, 1867 . Current status: Valid as Marcusenius ussheri (Günther, 1867) .

victoriae , Gnathonemus Worthington, 1929 . Current status: Valid as Marcusenius victoriae (Worthington, 1929) . [37 spp].