Orthomorpha scabra Jeekel, 1964

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2011, Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 131, pp. 1-161 : 47-49

Orthomorpha scabra Jeekel, 1964 Figs 4142

Pratinus granosus Attems 1953: 166 (D).

Orthomorpha granosa - Jeekel 1963: 265 (M).

Orthomorpha scabra Jeekel 1964: 361 (N, D).

Orthomorpha scabra - Jeekel 1968: 56 (M); Hoffman 1977: 700 (M); Golovatch 1998: 42 (M, D).

Lectotype.

♂ of Pratinus granosus (NHMW-3516), Vietnam, Lamdong Prov., Peak Lang Biang, 1938-1939, leg. C. Dawydoff.

Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male coming from a certain locality, because (1) Attems (1953) had provided no information on the number of syntypes, and (2) he stated their provenance to have been both from Xieng Kunang, Luang Prabang Province, Laos and Peak Lang Biang, Lamdong Province, Vietnam.

Redescription.

Length 41 mm, width of midbody pro- and metazona 3.1 and 4.4 mm, respectively (vs 38 mm in length and 4.6 mm in width, as given in the original description ( Attems 1953)). Coloration of alcohol material upon long-term preservation rather uniformly light brown (Fig. 41) (vs black-brown with rather contrasting light brown to yellow-brown prozona, paraterga and venter, as well as red-brown legs, as given in the original description ( Attems 1953)).

Head usual, clypeolabral region sparsely setose, surface of vertex smooth, epicranial suture very distinct. Antennae rather long and slender (Fig. 41A, B & J), extending behind body segment 3 dorsally. Head in width <collum <segments 3 and 4 <segment 2 <segments 5-17, gently and gradually tapering thereafter. Collum with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 2+2 posterior setae, all borne on small knobs; caudal corner of paraterga subrectangular, nearly pointed (Fig. 41A & B). Tegument dull; metaterga coriaceous, rugulose, each postcollum one with two rows of evident tubercles, caudal row being more evident: 2+2 and 3+3 on metatergum 2, 3+2 and 4+3 on metatergum 3, 2 –3+2– 4 until metatergum 17, 1+2 and 2+3 on metaterga 18 and 19; prozona very finely shagreened, surface below par aterga microgranulate. Tergal setae abraded. Axial line visible, starting from metatergum 2. Paraterga very strongly developed (Fig. 41A-G), mostly slightly upturned and lying above dorsum, lying below dorsum only on segments 2 and 3, thin in lateral view, like blunt blades, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, all pointed. Calluses on paraterga 2 delimited by a sulcus only dorsally, on following paraterga both dorsally and ventrally, rather broad, especially so on pore-bearing segments. Paraterga 2 broad, anterior edge rounded, lateral edge with two small incisions in anterior half; posterior edge evidently concave (Fig. 41A & B). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with one small incision in anterior half. Ozopores evident, lateral, not sunken inside a groove, lying at about 1/3 of metazonital length. Transverse sulcus complete on metaterga 4-18, incomplete on metatergum 3, shallow, reaching bases of paraterga, smooth at bottom, slightly sinuate anteromedially (Fig. 41A, B, C, D & F). Stricture between pro- and metazona rather narrow, shallow, slightly ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (Fig. 41B, D & E), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 11. Epiproct (Fig. 41E-G) conical, flattened dorsoventrally, apical papillae small; tip subtruncate; pre-apical papillae small, but visible, lying close to tip. Hypoproct (Fig. 41G) subtrapeziform, setiferous knobs at a subtruncate caudal margin small and well-separated.

Sterna densely setose, without modifications, but with two large, rounded, fully separated, setose cones between ♂ coxae 4 (Fig. 41H & I). A paramedian pair of conspicuous, high tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3-1.4 as long as body height, prefemora without modifications, tarsal brushes present until ♂ legs 5.

Gonopods (Fig. 42) simple. Coxa long and slender, with several setae distodorsally. Prefemur densely setose, about 2 times shorter than femorite + “postfemoral” part. Femorite very slender, evidently curved, slightly enlarged distad, “postfemoral” part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two subequal denticles (one terminal, the other middle) and a broader subterminal lobule.

Remark.

This species was renamed to avoid homonymy ( Jeekel 1964).