Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes & Nguyen 2015

Phimmachak, Somphouthone, Aowphol, Anchalee & Stuart, Bryan L., 2015, Morphological and molecular variation in Tylototriton (Caudata: Salamandridae) in Laos, with description of a new species, Zootaxa 4006 (2), pp. 285-310 : 297

Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes & Nguyen 2015

Tylototriton shanjing Group 1 Nishikawa, Khonsue, Pomchote & Matsui 2013a: 262.

Referred material. NCSM 82952–53, NUOL 0 0 420 (three males), NUOL 0 0 419 (one female), NUOL 0 0 448 (one eft), Laos, Luang Namtha Province, Viengphoukha District, Phou Ya Khao Village, Nam Long Stream, 20.89236ºN 101.08519ºE, 1,466 m elev., coll. 8 July 2011 by Somphouthone Phimmachak and Sengvilay Seateun ( Figs. 4 C–H).

Measurements are summarized in Table 5 .

Distribution and natural history. Specimens were collected during the day under grass in shrub forest 2–6 m from a 5–7 m wide stream. The shrub forest had been burned within the previous five years.

This is the first report of the species in Laos, although the type locality at Muong Nhe, Dien Bien Province, Vietnam is very near to the border of Laos, and so it was expected there ( Le et al. 2015). Elsewhere the species is known from Thuan Chau, Son Lao Province, Vietnam, and Doi Lahnga, Chiang Rai Province, Thailand ( Le et al. 2015; Fig. 2 ).

Figures

FIGURE 2. Principal component analysis of the genus Tylototriton showing two dimensional plots of the first principal component (PC 1) versus the second (PC 2). (A) males of the subgenus Tylototriton, (B) females of the subgenus Tylotriton, and (C) males of the subgenus Yaotriton.

FIGURE 4. Tylototriton notialis: (A) Dorsal view and (B) ventral view of topotype female (NUOL 00443) at Phou Ak, Nakai Nam Theun National Protected Area, Boualapha District, Khammouan Province, Laos. Tylototriton anguliceps: (C) Dorsal view of female (NUOL 00419), (D) dorsal view of male (NUOL 00420), (E) ventral views of female above (NUOL 00419) and male below (NCSM 82953), (F) dorsal view of eft (NUOL 00448), (G) microhabitat at stream bank, and (H) macrohabitat at Nam Long Stream, Phou Ya Khao Village, Viengphoukha District, Laos, Luang Namtha Province, Laos.