Thymioptila clavata, Abstract, 2022

Abstract, Nantasak Pinkaew, 2022, A new species of genus Thymioptila Diakonoff (Lepidoptera: Tortricidae) from Thailand and a new record, Thymiotila oedalea (Meyrick, 1909), from Laos, Zootaxa 5222 (6), pp. 534-544 : 535-536

Thymioptila clavata sp. nov.

urn:lsid:zoobank.org:act:515582A3-9CC1-4C19-A932-45E7288FA4E0

( Figs. 1–2 , 5 , 8–10 , 11 , 13 , 15 )

Type Material. Holotype, ♁. THAILAND; Sa Kaeo Prov., Pang Sida N.P., 14 o 00ʹ54ʺN, 102 o 11ʹ45ʺE, alt. 260 m, 27 Feb. 2017, Pinkaew et al. leg., np9443 (genitalia slide NP3962); KKIC.

Paratypes: 2♁. THAILAND; Chanthaburi Prov., Khao Soi Dao N.P., 13o05ʹ49ʺN, 102o10ʹ21ʺE, alt. 166 m, 17 Mar. 2008, np2752 (genitalia slide NP1532) ; Chaiyaphum Prov., Phu Khieo W.S., 16o30ʹ24ʺN, 101o41ʹ22ʺE, alt. 770 m, 16 Mar. 2018, np11987 (genitalia slide NP3976). All specimens were collected by Pinkaew et al. and deposited in KKIC .

Diagnosis. This new species is similar to Thymioptila oedalea (Meyrick) . Forewing of both species have a tuft of the longitudinal raised scales at posterior end of closed cell. These scales are quite distinct and more raised up in resting posture of living specimens ( Fig. 1 ). Labial palpus of both species has similar pattern ( Figs. 5–7 ). Marking at the middle of costa is slightly different by a moderately large subtriangular mark in T. oedalea ( Figs. 3–4 ) but a rather small spot in Thymioptila clavata sp. nov. ( Fig. 2 ). Hind wing of T. clavata sp. nov. has a tuft of long hair pencils on anal marginal groove on the upper side ( Fig. 9 ) and a row of long setaform scales along 3A on the underside ( Fig. 10 ), but both characters are absent in T. oedalea . Hind tibia in male of T. clavata sp. nov. is enlarged by covered bushy scales ( Fig. 11 ) but hind tibia of T. oedalea is simple and covered with long and narrow setaform scales ( Fig. 12 ). Pregenital abdomen of both species can be separated by shape of sclerotized plate on tergum VIII (broad subtriangular in T. clavata sp. nov. but elongate subtriangular in T. oedalea ) and sternum VIII (narrow band connected with slightly sinuate band in T. clavata sp. nov. but narrow band connected with strongly curve band in T. oedalea ), especially two large patches of dense scale sockets between tergum VII and sternum VII on posterior 1/ 2 in T. clavata sp. nov. but absent in T. oedalea ( Figs. 13–14 ). Male genitalia of T. clavata sp. nov. can be distinguished by subtriangular uncus, subtriangular cucullus with a large curved spine on large subtriangular lobe at apex and a large claviform cucullus process ( Fig. 15 ) whereas uncus is sub-semicircular, cucullus is narrow and curve with a large curved spine at apex and quite narrow claviform cucullus process in T. oedalea ( Fig. 16 ).

Description. Head ( Fig. 5 ). Lower frons yellowish white; upper frons light brown; vertex light brown mixed with brown near chaetosema, posterior margin of compound eyes edged with brown alternated with white scales; antenna light brown; labial palpus with first segment white, second segment enlarged, parallel margins except base, basal half white, apical half light brown with a large transverse brown patch apically, from dorsal to ventral margin, third segment short, brown.

Thorax: pronotal collar light brown mixed with some purple-brown scales, tegulae light brown ground with large patch inner half, brown mixed with dark brown; mesonotum light brown mixed with greyish brown, without posterior crest; Hind legs ( Fig. 11 ) strongly modified, inner side of femur with patch of dense, long white scales along ventral margin, tibia enlarged by dense scales, slightly narrowed at posterior end, mostly white mixed with dark brown beneath. Forewing subrectangular, wingspan 19.5–20.0 mm in males ( Fig. 2 ) (n = 3); costal margin rather straight, gently curved at apical 1/4, termen truncate, rather straight from wing apex to M 3 then curved to tornus; costal strigulae well developed, creamy white separated by small dark brown streaks, 1 st –2 nd indistinct, 3 rd –4 th double pairs, 5 th –9 th more distinct pair on apical half; ground color creamy white mixed with scattered spots and short, light brown longitudinal lines throughout, basal 1/5 with rather small indistinct patch, brown mixed with dark brown, middle of costa with a small oblique mark, brown mixed with dark brown extending from costa to R 1, apical 1/3 with a large indistinct transverse patch, brown mixed with dark brown, extending from R 4 to tornus broken by a narrow, longitudinal raised scale patch at the posterior end of closed cell between base of R 5 and M 2 ( Fig. 8 ), more raised up in living specimens ( Fig. 1 ); fringe scales light brown; underside brown with narrow creamy strip along costa apical 1/2. Hindwing ( Figs. 8–10 ) brown slight paler to base, anal area with long hairy scales, light brown, anal margin sclerotized, raised upward forming longitudinal groove, covered with white scales, bearing a tuft of long hair pencils, yellowish white; underside brown, slightly paler to base, except apical 1/2 of costa with a narrow creamy white strip and brownish yellow anal area with a distinct row of light brown long setaform scales along 3A medially.

Abdomen. Pregenital abdomen ( Fig. 13 ) with a narrowed sclerotized T-shaped band on sternum VIII; tergum VIII with a moderately sclerotized subtriangular plate, slightly tapered posteriorly to rounded apex, with two patch of dense scales socket on membrane between tergum VII and sternum VII on posterior 1/2. Male genitalia ( Fig. 15 ) with tegumen subtriangular, moderately wide, apicolateral 1/2 with dense scale sockets, basal 1/2 with sparsely scale sockets; uncus moderately large, subtriangular, rounded apex, with long, sparsely setose, posterior surface with two separated patch of dense scale sockets; socii narrow, membranous, densely setose; gnathos arising from near midlength of tegumen, forming an upcurved sclerotized narrow band; vinculum rather narrow; juxta subtriangular; caulis short; anellus a high hood-like surrounding basal half of phallus; phallus moderately long and wide, slightly sclerotized, margins rather parallel; tuba analis with wrinkled surface; cornuti absent; valva rather long and slender; sacculus with widened basal opening, basal half sparsely setose, apical half with moderately dense setae, costal margin with moderately dense setae at basal 1/2; neck distinct with deep incised ventrally; cucullus short forming moderately large subtriangular lobe, pointing ventrally with a large, pointed, curved spine at apex, covered with moderately dense scale sockets, with sparse long setae ventrally, dorsoapical corner with a distinct large and long clavate process, rounded apex, slightly curved upward and protruding outward, strongly widened apical 1/2, basal 1/2 with moderately dense scale sockets, apical 1/2 densely setose.

Female. Unknown.

Distribution. Thailand (Sa Kaeo, Chanthaburi, and Chaiyaphum Province).

Etymology. The specific epithet ( clavata = clavate) refers to the shape of the cucullus process.

Figures

FIGURE 1. Living male specimen of Thymioptila clavata sp. nov. (paratype, np 11987) with natural resting posture.

FIGURES 2–4. Wing pattern of Thymioptila spp. 2. T. clavata sp. nov. (holotype). 3. T. oedalea (male). 4. T. oedalea (female). Scale bars = 2 mm.

FIGURE 8. A tuft of raised scales on forewing of Thymioptila clavata sp. nov. (holotype).

FIGURES 9–10. Modified scales on anal area of hindwing of Thymioptila clavata sp. nov. (holotype). 9. A tuft of hair pencils tuft in anal fold and dense of long scales along 3A (upper side). 10. A row of long hair pencils along 3A (underside).

FIGURES 5–7. Head of Thymioptila spp. 5. T. clavata sp. nov. (holotype). 6. T. oedalea (male). 7. T. oedalea (female).

FIGURES 11–12. Sex scales on hind leg of Thymioptila spp. 11. T. clavata sp. nov. (holotype). 12. T. oedalea.

FIGURES 13–14. Sclerotized plates on 8th tergum (left) and sternum (right) of Thymioptila spp. 13. T. clavata sp. nov. (holotype). 14. T. oedalea.

FIGURES 15–16. Male genitalia of Thymioptila spp. 15. T. clavata sp. nov. (holotype). 16. T. oedalea.