APHODIINAE, Arrow, 1909

Peck, Stewart B., 2016, The beetles of the Lesser Antilles (Insecta, Coleoptera): diversity and distributions, Insecta Mundi 2016 (460), pp. 1-360 : 83-87

SUBFAMILY APHODIINAE

TRIBEAPHODIINI

Labarrus lividus (Olivier) 1789: 86 ( Scarabaeus ); Arrow 1903 502 ( Aphodius ); Chapin 1940a: 6; Paulian 1947a: 36 ( Nialus ); Chalumeauand Gruner 1974: 794; Cartwrightand Chalumeau 1978: 7; Chalumeau 1983a: 57 ( Aphodius ( Nialus )); Valentine and Ivie 2005: 276; Gordon and Skelley 2007: 262, generic synonymy; Skelley et al. 2007: 5. Distribution. Antigua *!, Barbados *!, Cuba, Désirade, Dominica, Grenada!, Guadeloupe, Guana, Hispaniola, Jamaica, Les Saintes, Marie-Galante, Martinique, Montserrat*!, Nevis *!, Puerto Rico, Saba*!, St. Croix, St. Kitts *!, St. Lucia *!; perhaps on practically all West Indian islands. Nearly cosmopolitan; probably from USA to Mexico to Argentina; native to the Old World and introduced to the New World; introduced to the Lesser Antilles. Notes. This species has commonly been confused with Labarrus pseudolividus and many island records listed above (mostly from Chapin 1940a: 6) need confirmation (Gordon and Skelley 2007: 263). An exclamation mark (!) indicates islands for which we have seen voucher material. Common in cow dung and other excrement.

Labarrus pseudolividus (Balthasar) 1941: 148 ( Aphodius ); Marquet and Roguet 2003: 11; Gordon and Skelley 2007: 263; Skelley et al. 2007: 5; Ivie et al. 2008b: 244 ( Aphodius ); Turnbowand Thomas 2008: 49; Thomaset al. 2013: 42. Distribution. Bahamas, Caymans, Dominica *!, Martinique, Montserrat!, Nevis *!, St. Vincent *!; introduced to the Lesser Antilles. Introduced and now widespread in the New World; probably on most of the islands listed for Labarrus lividus above, with which it has been commonly confused. Notes. Previously cited material must be re-examined to accurately delimit the distribution of this species. An exclamation mark (!) indicates islands for which we have seen voucher material.

Nialaphodius nigrita (Fabricius) 1801: 73 ( Aphodius ); Marquet and Roguet 2003: 11; Gordon and Skelley 2007: 265; Ivieet al. 2008b: 244 ( Aphodius ); Daltry 2009: 65; Thomaset al. 2013: 43. = Aphodius cuniculus Chevrolat 1864b: 411 , Chapin 1940a: 7; Ramos 1946: 41; Wolcott 1951: 248 of PuertoRico, of Mona, of Vieques; Bordat 1990: 62, synonymy; Paulian 1947a: 37 ( Nialus ); Chalumeauand Gruner 1974: 795; Cartwrightand Chalumeau 1978: 8; Bennettand Alam 1985: 22; Woodruff et al. 1998: 32; Chalumeau 1983a: 58 ( Aphodius ( Nialus )); Marquet and Roguet 2003: 11; Valentine and Ivie 2005: 276; Skelley et al. 2007, generic synonymy; Turnbow and Thomas 2008: 49; Daltry 2009: 65 of St. Lucia. = Aphodius granarius variety guadeloupensis Fleutiauxand Sallé 1890: 395 . Distribution. Antigua, Barbados, Carriacou, Caymans, Cuba, Désirade, Dominica, Grenada, Guadeloupe, Guana, Hispaniola, Les Saintes, Jamaica, Marie-Galante, Martinique, Mona, Montserrat, Mustique, Nevis *, Puerto Rico, Saba*, St. Croix, St. John, St. Kitts, St. Lucia, St. Thomas, St. Vincent, Tortola, Vieques. Widespread southern USA, Mexico to widespread South America , Tobago; introduced to the Lesser Antilles. Introduced to New World from Afro-tropical region; worldwide in tropics and subtropics. Notes. Common in cow dung. Plate 11.

TRIBEEUPARIINI

Chapin (1940a) is an out of date summary of the West Indian fauna. Stebnicka (2007b) is an illustrated and up-to-date summary of the New World fauna of the very large genus Ataenius , and has provided an extensive review of the New World fauna (see citations below), but some of her synonymies of Lesser Antillean species names are not accepted by F. Chalumeau (pers. comm., December 2010) who has extensive experience with this group, and these are noted below.

Ataeniopsis armasi (Chalumeau) 1982a: 321 ( Ataenius ); Stebnicka 2003b: 109. = Ataeniopsis haroldi (Steinheil) 1872: 556 ( Ataenius ) of Guadeloupe, Chalumeauand Gruner 1974: 802 (misidentification); Chalumeau 1983a: 69 ( Ataenius ); Stebnicka 2003b: 104, 2009: 54. Distribution. Guadeloupe, Hispaniola, Puerto Rico; widespread Antillesendemic. Notes. Chalumeau (in litt., Nov., 2012) thinks the record of Guadeloupe of A. haroldi (Steinheil) is valid and that A. armasi is not of Guadeloupe (Chalumeau 1983a: 69) but is restricted to the Greater Antilles. The distributions of both species still need clarification.

Ataenius aequalis Harold 1880: 40 ; Stebnicka 2005: 103; Stebnicka 2007b: 44. = Ataenius insulicola Chapin 1940a: 28 of St. Vincent (type locality). Distribution. Guadeloupe, St. Vincent. USA (LA), Mexico to Trinidad to Argentina; widespread New World. Notes. Both names are good species and neither are of Guadeloupe (Chalumeau, in litt., Nov., 2012).

Ataenius arenosus Harold 1868: 86 ; Stebnicka 2007b: 77, P. Skelley det., new species record. Distribution. St. Vincent *. Venezuela, Surinam, Guyana, Brazil; the Lesser Antillesand Latin America .

Ataenius attenuator Harold 1874: 2 ; Stebnicka 2007b: 68; Stebnicka 2007b: 68. = Ataenius abditoides Chapin 1940a: 18 of St. Lucia. Distribution. Grenada *, St. Lucia, St. Vincent. Mexico to Panama toTrinidad, Brazil, and Peru; the Lesser Antillesand Latin America .

Ataenius beattyi Chapin 1940a: 17 ; Blackwelder 1944-1957: 213; Ramos 1946: 41; Bennett and Alam 1985: 22; Valentineand Ivie 2005: 276; Stebnicka 2006: 107; Stebnicka 2007b: 57; Daltry 2009: 65; Thomas et al. 2013: 42. = Ataenius cameneni Chalumeau and Gruner 1974: 801 ; Chalumeau 1983a: 67; Ivieet al. 2008b: 244. Distribution. Antigua*, Barbados, Caymans, Désirade, Grenada *, Guadeloupe, Guana, Hispaniola, Jamaica, Mayreau*, Mona, Montserrat, Nevis*, PuertoRico, St. Croix , St. Kitts*, St. Lucia, St. Thomas, St. Vincent*, Union *; widespread Antilles endemic. Notes. In fowl dung on Barbados. F. Chalumeau (pers. comm., December 2010) does not agree with Stebnicka’s placement of Ataenius cameneni as a junior synonym, but views it as a distinct species. Plate 9 .

Ataenius brevicollis (Wollaston) 1854: 229 ( Oxyomus ); Chalumeau and Gruner 1974: 807; Chalumeau 1978: 42, 1983a: 77; Stebnicka 2007a: 58, 2007b: 63; Ivieet al. 2008b: 244; Turnbowand Thomas 2008: 48; Thomas et al. 2013: 42. = Ataenius sulcatulus Chevrolat 1864b: 413 of Cuba ; Fleutiaux and Sallé 1890: 397 of Guadeloupe ; Paulian 1947a: 42 of Guadeloupe. = Ataenius lherminieri Paulian 1947a: 43 of Guadeloupe ; Chalumeau 1983a: 77. Distribution. Antigua*, Bahamas, Barbados *, Caymans, Cuba, Désirade , Guadeloupe , Hispaniola , Jamaica, Les Saintes , Martinique, Montserrat. USA (FL, MS, TX) ; widespread Antilles and North and/or Central America . Introduced to Madeira, Canary Islands, Portugal ; anthropophilic. Notes. The synonymy of Ataenius lherminieri is in doubt, and the species has not been seen since the type collection ( Chalumeau 1983a: 78). Taken on bat guano in a cave as well as at light traps .

Ataenius carinator Harold 1874: 20 ; Stebnicka 2007b: 76; Turnbowand Thomas 2008: 48; Thomas et al. 2013: 42. = Ataenius vincentiae Arrow 1903: 513 ; Chapin 1940a: 19; Chalumeauand Gruner 1974: 801; Cartwright and Chalumeau 1978: 11; Chalumeau 1983a: 65; Marquet and Roguet 2003: 11; Ivie et al. 2008b: 244; Schiller 2004: 42. Distribution. Antigua *, Bahamas, Caymans, Dominica, Guadeloupe, Hispaniola, Martinique, Montserrat, Nevis *, Puerto Rico, St. Lucia * (also in Daltry 2009: 65), St. Vincent. USA (FL), Mexico to Costa Rica, Venezuela to Bolivia, Brazil, Peru; widespread New World. Notes. Adults attracted to lights; found in cow dung. F. Chalumeau (in litt., Nov., 2012) does not agree with Stebnicka’s placement of Ataenius vincentiae as a junior synonym, but views it as a distinct species.

Ataenius cartwrighti Chalumeau and Gruner 1974: 813 ; Chalumeau 1983a: 86; Stebnicka and Lago 2005: 66; Stebnicka 2007b: 39. Distribution. Grenada *, Guadeloupe; Lesser Antillesendemic.

Ataenius communis Hinton 1936: 421 ; Stebnicka 2007b: 16; det. P. Skelley, new species record. Distribution. St. Vincent *. El Salvador to Panama, Colombia, Venezuela, Trinidad; the Lesser Antilles and Latin America .

Ataenius complicatus Harold 1869: 102 ; Stebnicka 2007b: 53; det. P. Skelley, new species record. Distribution. Grenada *. Mexico to Panama, Colombia to Bolivia to Venezuela and Netherland Antilles; the Lesser Antillesand Latin America .

Ataenius crenulatus Schmidt 1910: 359 ; Stebnicka 2006: 108; Stebnicka 2007b: 57. Distribution. Barbados; introduced. Mexico , Honduras, Venezuela, Brazil, Bolivia, to Argentina; introduced to the Lesser Antilles.

Ataenius cribrithorax Bates 1887: 95 ; Chapin 1940a: 28; Cartwrightand Chalumeau 1978: 14; Stebnicka 2007b: 16. Distribution. Cuba, Dominica, Jamaica, Martinique, St. Thomas. Mexico to Panama; widespread Antilles and North and/or Central America . Notes. Adults attracted to lights; found in cow dung. The possibilityneeds tobechecked that this is a synonym of Ataenius nugator (F. Chalumeau, in litt., December, 2010).

Ataenius gracilis (Melsheimer) 1845b: 137 ( Oxyomus ); Fleutiaux and Sallé 1890: 397; Chapin 1940a: 25; Paulian 1947a: 44; Chalumeauand Gruner 1974: 806; Cartwrightand Chalumeau 1978: 12; Chalumeau 1978: 44, 1983a: 74; Bennett and Alam 1985: 22; Woodruff et al. 1998: 33; Marquet and Roguet 2003: 11; Stebnicka 2007b: 79; Ivieet al. 2008b: 244; Turnbowand Thomas 2008: 48; Thomaset al. 2013: 42. = Ataeniuschilensis Solier 1851: 72 ; Woodruff et al. 1998: 33, of Barbados. Distribution. Antigua, Bahamas, Barbados, Caymans, Cuba, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Marie- Galante, Martinique, Montserrat, PuertoRico, Saba*, St. Croix, St. Kitts, St. Lucia *, St. Vincent, Vieques. Canada, United States; throughout the Neotropics ( Argentina, Chile, Colombia, Peru, Galapagos Islands); widespread New World. Notes. Adults attracted to lights; found in cow dung. Plate 9.

Ataenius heinekeni (Wollaston) 1854: 228 ( Oxyomus ); Stebnicka 2004: 216 (synonymy); Thomas et al. 2013: 42. = Ataenius rhyticephalus (Chevrolat) 1864: 413 ( Auperia ); Bennettand Alam 1985: 22; Turnbowand Thomas 2008: 48. Distribution. Bahamas, Barbados (probably introduced), Caymans, Cuba, Hispaniola, Puerto Rico, St. Thomas. USA (SC-FL-TX), Mexico to Brazil, introduced to Ascension and Madeira islands; introduced to the Lesser Antilles? Notes. Taken by beating, in leaf and log litter and in debris; seemingly not at dung.

Ataenius howdeni Chalumeau 1978: 51 ; Stebnicka 2002: 269; Stebnicka 2007b: 21; Ivie et al. 2008b: 244. Distribution. Antigua, Montserrat; Lesser Antillesendemic.

Ataenius hygrophilus Paulian 1947a: 39 ; Chalumeau and Gruner 1974: 809; Chalumeau 1983a: 79; Stebnicka 2007a: 57; Stebnicka 2007b: 62. Distribution. Guadeloupe; single island endemic. Notes. The species has not been found again (Chalumeau 1983a: 80). Plate 9.

Ataenius imbricatus (Melsheimer) 1845b: 136 ; Stebnicka 2003a: 225; Stebnicka 2007b: 27; Turnbow and Thomas 2008: 48; Thomaset al. 2013: 42. Distribution. Bahamas, Caymans, Cuba, Hispaniola, Puerto Rico, St. Croix, Guadeloupe, Barbados. Mexico , Central America , Colombia to Trinidad to Brazil and Argentina; widespread Antilles and Latin America . Notes. In cattle dung; mostly collected at lights. Chalumeau (in litt., Nov., 2012) does not accept that the species is of Guadeloupe so the record would then be a misidentification.

Ataenius insulae Chalumeau and Gruner 1974: 805 ; Chalumeau 1983: 72; Stebnicka 2002: 275; Stebnicka 2007b: 24. Distribution. Guadeloupe; singleisland endemic.

Ataenius liogaster Bates 1887: 94 ; Chapin 1940a: 29; Paulian 1947a: 44; Chalumeau and Gruner 1974: 813; Stebnicka and Lago 2005: 60 (synonymy); Stebnicka 2007b: 36; Ivieet al. 2008b: 244; Thomas et al. 2013: 42. = Ataenius edwardsi Chapin 1940a: 26 ; Cartwright and Chalumeau 1978: 14; Chalumeau 1983a: 82, 174; Bennett and Alam 1985: 22; Woodruff et al. 1998: 33; Stebnicka and Lago 2005: 60. Distribution. Antigua, Barbados, Carriacou, Caymans, Cuba, Dominica (overlooked in Stebnicka and Lago 2005: 61), Grenada, Guadeloupe, Hispaniola, Jamaica, Montserrat, Puerto Rico, St. Croix, St. Lucia, St. Vincent, Union*. Mexico to Panama, Ecuador (including Galapagos), Trinidad, Venezuela; Micronesia; widespread Antilles and Latin America . Notes. Adults attracted to lights; found in cow dung.

Ataenius luteomargo Chapin 1940a: 36 ; Paulian 1947a: 41; Chalumeauand Gruner 1974: 811; Cartwright and Chalumeau 1978: 12; Chalumeau 1983a: 84; Bennettand Alam 1985: 22; Woodruff et al. 1998: 33; Stebnicka 2002: 270, 2007b: 22. = Ataenius terminalis Fleutiauxand Sallé 1890: 396 ; Uyttenboogaart 1902: 116, of Barbados; Arrow 1903: 512 (not Chevrolat). = Ataenius versicolor Hinton 1937a: 183 (not Schmidt), of Barbados. Distribution. Antigua, Barbados, Désirade, Dominica (type locality), Grenada, Guadeloupe, Hispaniola, Les Saintes, Marie-Galante, Martinique, Montserrat, Puerto Rico, St. Kitts, St. Lucia ; not Jamaica. Panama, Surinam, Venezuela; widespread Antillesand Latin America . Notes. Adults attracted to lights, found in cow dung. Plate 9.

Ataenius morator Harold 1869: 103 ; Stebnicka 2003a: 243, 2007b: 32. = Ataenius picipes Fleutiaux and Sallé 1890: 397 ; Paulian 1947a: 43; Chalumeauand Gruner 1974: 804; Cartwrightand Chalumeau 1978: 13; Chalumeau 1978: 44, 1983: 71, 1992: 200 (synonymy); Touroult 2005: 84. = Ataenius tenebrosus Arrow 1903: 512 ; Chapin 1940a: 23; Chalumeauand Gruner 1974: 804 (synonym of Ataenius picipes ). Distribution. Antigua *, Barbados, Dominica, Grenada, Guadeloupe, Hispaniola, Martinique, Mayreau*, Montserrat*, Mustique*, Puerto Rico, St. Lucia, St. Vincent *, Vieques. Trinidad, Venezuela south to Argentina; widespread Antilles and Latin America . Notes. In rotting materials, especially dung; frequent in light traps.

Ataenius nugator Harold 1880: 41 ; Chalumeau and Gruner 1974: 812; Chalumeau 1983a: 83; Marquet and Roguet 2003: 11; Stebnicka 2007b: 15. Distribution. Cuba, Dominica, Jamaica, Martinique, St. Thomas. Mexico to Panama, Colombia and Venezuela to Brazil, Bolivia, Peru; widespread Antilles and Latin America . Notes. Adults attracted to lights; found in cow dung.

[ Ataenius platensis (Blanchard) 184737-1846: 185 ( Oxyomus ); Stebnicka 2005: 124, 2007b: 50; Turnbow and Thomas 2008: 48. Distribution. Bahamas, Mexico to Venezuela to Chile and Argentina; widespread Greater Antilles (not the Lesser Antilles). Not Guadeloupe, not Martinique; the records of Hinton 1937a: 177 are questioned by Chalumeau 1983a: 215.]

Ataenius picinus Harold 1867: 281 ; Chalumeauand Gruner 1974: 810; Cartwrightand Chalumeau 1978: 14; Chalumeau 1978: 44, 1983a: 80; Marquet and Roguet 2003: 11; Stebnicka 2004: 224, 2007b: 42. = Ataenius elongatus Palisot de Beauvois 1811: 104 ; Fleutiaux and Sallé 1890: 397 of Guadeloupe. = Ataenius darlingtoni Hinton 1937a: 179 ; Chapin 1940a: 30; Ramos 1946: 41; Paulian 1947a: 40. Distribution. Antigua, Cuba, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Marie-Galante, Martinique, Mayreau*, Mona, Montserrat*, Puerto Rico, St. Croix, St. Vincent. United States, Argentina, Brazil, Uruguay; widespread New World; Australia, Fiji, New Caledonia, New Hebrides, New Zealand; nearly cosmopolitan. The most widely distributed species in the genus, and undoubtedly on all islands of the Lesser Antilles. Notes. Adults attracted to lights; found in cow dung. Plate 9.

[ Ataenius polyglyptus Bates 1887: 99 ; Stebnicka 2007b: 73. Distribution. Ecuador, Guatemala, Panama. Not Mustique, not Grenada, not St. Vincent; errors in Blackwelder 1944-1957: 215.]

Ataenius punctipennis Harold 1868: 86 ; Stebnicka 2005: 130; Stebnicka 2007b: 48. = Ataenius subopacus Chapin 1940a: 24 . Distribution. Bequia*, Canouan*, Carriacou, Grenada, Mayreau*, Mustique*, Union*. Colombia, Trinidad, Venezuela, Brazil, Argentina; the Lesser Antillesand Latin America .

Ataenius scabrellus Schmidt 1909: 118 ; Chalumeau and Gruner 1974: 807; Stebnicka 2003: 227, 2007b: 27; Ivieet al. 2008b: 244; Turnbowand Thomas 2008: 48; Thomaset al. 2013: 42. = Ataenius havanensis Balthasar 1938: 56 ; Chalumeauand Gruner 1974: 807; Chalumeau 1983a: 75. = Ataenius miamii Chapin 1940a: 41 ; Ramos 1946: 41; Bennettand Alam 1985: 22; Tucker 1952: 342 of Barbados; Miskimen and Bond 1970: 97 of St. Croix; this is a valid species and it is not known from the Lesser Antilles (Stebnicka 2007b: 27). = Ataenius fleutiauxi Paulian 1947a: 42 of Guadeloupe. Distribution. Bahamas, Barbados, Caymans, Cuba, Bahamas, Désirade, Hispaniola, Jamaica, Les Saintes, PuertoRico, Grenada *, Guadeloupe, Martinique, Mona, Montserrat, Mustique*, St. Croix, St. Kitts. USA (FL); Venezuela, Guyana; widespread Antillesand Latin America . Notes. Fortuné Chalumeau (in litt., Nov., 2012) indicates that Stebnicka’s placement of Ataenius havanensis as a synonym is wrong and that it is a valid species as is A. miamii but they are not of Guadeloupe. Plate 9.

Ataenius sculptilis Harold 1868: 86 ; Stebnicka 2006: 102; Stebnicka 2007b: 56; det. P. Skelley, 2011, new species record. Distribution. Grenada *, Puerto Rico, St. Thomas. Trinidad, Venezuela and Brazil to Peru and Argentina; widespread New World.

Ataenius scutellaris Harold 1867: 82 ; Chalumeau and Gruner 1974: 803; Cartwrightand Chalumeau 1978: 14; Chalumeau 1978: 44, 1983a: 69; Bennett and Alam 1985: 22; Marquet and Roguet 2003: 11; Valentineand Ivie 2005: 276; Stebnicka 2007a: 48, 2007b: 59; Ivieet al. 2008b: 244. = Ataenius frater Arrow 1903: 512 of St. Vincent; Chapin 1940a: 32; Woodruff et al. 1998: 33 of Grenada. = Ataenius elongatus Palisot de Beauvois 1811: 104 ; Fleutiaux and Sallé 1890: 397 of Guadeloupe, misidentification. Distribution. Antigua, Bahamas, Barbados, Cuba, Dominica, Grenada, Guadeloupe, Guana, Hispaniola, Jamaica, Marie-Galante, Martinique, Montserrat, PuertoRico, Saba*, St. Croix, St. Kitts, St. Lucia, St. Martin-St. Maarten, St. Thomas, St. Vincent, Tortola. Mexico to Bolivia and Brazil, Trinidad; widespread Antilles and Latin America ; introduced to Vanuatu ( New Hebrides), Seychelles, Mascarenes, Malaya. Notes. Adults attracted to lights; found in cow dung.

Ataenius steinheili Harold 1874: 18 ; Chapin 1940a: 16; Chalumeau 1978: 43; Stebnicka 2006: 105; Stebnicka 2007b: 57. = Ataenius grenadensis Chalumeau 1982a: 327 of Grenada. Distribution. Grenada. Mexico , Colombia, Trinidad, Venezuela; the Lesser Antillesand Latin America .

Ataenius strigicaudus Bates 1887: 96 ; Chapin 1940a: 32; Paulian 1947a: 40; Chalumeau and Gruner 1974: 810; Cartwrightand Chalumeau 1978: 13; Chalumeau 1983a: 81; Bennettand Alam 1985: 22; Woodruff et al. 1998: 33; Marquet and Roguet 2003: 11; Stebnicka 2004: 214, 2007b: 40. Distribution. Bahamas, Barbados, Bequia, Cuba, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Martinique, Montserrat, Puerto Rico, St. Croix, St. Lucia, St. Thomas, St. Vincent. Mexico to Trinidad, Chile and Argentina; widespread Antilles and Latin America . Notes. Adults attracted to lights; found in cow dung. Plate 9.

Ataenius temperei Chalumeau and Gruner 1974: 799 ; Cartwright and Chalumeau 1978: 12; Chalumeau 1983a: 64; Stebnicka 2007b: 69; Ivieet al. 2008b: 244; Touroult 2005: 84. Distribution. Dominica, Guadeloupe (type locality), Montserrat, St. Kitts ; Lesser Antilles endemic. Notes. Adults attracted to lights; not found on cow dung.

Euparia baraudi Chalumeau and Gruner 1974: 796 ; Cartwrightand Chalumeau 1978: 10; Chalumeau 1983a: 59; Chalumeauand Howden 1984: 88; Stebnicka 2009: 14. Distribution. Dominica, Grenada *, Guadeloupe (type locality) ; Lesser Antilles endemic. Records of Grenada and of Trinidad are not verified in Stebnicka 2009: 14. Notes. Caught at lights, possible myrmecophile. Plate 10.

Iguazua blackwelderi (Chapin) 1940: 11 ( Saprosites ); Ivie et al. 2008b: 244; Stebnicka 2009: 51. Distribution. Barbados *, Guadeloupe, Montserrat, St. Lucia, Puerto Rico. Guyana; the Lesser Antilles and Latin America . Notes. Collected in decayed wood, rotten logs, and under bark of dead trees.

Saprosites dufaui Paulian 1947a: 46 ; Chalumeau and Gruner 1974: 815; Chalumeau 1983a: 88; Marquet and Roguet 2003: 12; Stebnicka 2009: 46. Distribution. Guadeloupe, Martinique; Lesser Antilles endemic. Notes. Found under bark at 600 m. The Martinique record needs confirmation. Plate 13

Saprosites exaratus (Fleutiaux and Sallé) 1890: 397 ( Ataenius ); Chalumeau and Gruner 1974: 815; Chalumeau 1983a: 89; Touroult 2005: 84; Stebnicka 2009: 46; Daltry 2009: 65. = Saprosites wirthi Cartwrightand Chalumeau in Chalumeau 1977a: 72; Cartwrightand Chalumeau 1978: 16; Chalumeau 1983a: 90; Marquet and Roguet 2003: 12; Stebnicka 2009: 46 (synonymy). Distribution. Dominica, Guadeloupe, Martinique, St. Lucia; Lesser Antillesendemic. Notes. Chalumeau (in litt., Nov., 2012) does not accept the synonymy of S. wirthi Cartwright and Chalumeau of Dominica into this species. Plate 13.

Saprosites grenadensis Arrow 1903: 574 ; Chapin 1940a: 11; Stebnicka 2009: 47. Distribution. Grenada, Martinique, St. Lucia *. Trinidad, Venezuela; the Lesser Antillesand Latin America . Notes. Adults and immatures taken in decaying heart of Euterpe sp. palm. Plate 13.

TRIBEPSAMMODIINI

Platytomus parvulus (Chevrolat) 1864: 415 ( Psammodius ); Paulian 1947a: 47 ( Diastictus ); Chapin 1940a: 8; Chalumeauand Gruner 1974: 815; Cartwrightand Chalumeau 1978: 15; Chalumeau 1983a: 91; Woodruff et al. 1998: 34; Marquet and Roguet 2003: 12; Thomaset al. 2013: 43. Distribution. Antigua *, Barbados *, Caymans, Cuba, Dominica, Grenada, Guadeloupe, Jamaica, Martinique, St. Croix, St. Lucia *, St. Vincent, Union*. Colombia, Trinidad; widespread Antillesand Latin America . Notes. Ahumus feeder in soil, flying at dusk and attracted to lights. Plate 12.

Psammodius cameneni Chalumeau 1976: 128 ; Cartwright and Chalumeau 1978: 15; Chalumeau 1983a: 92. Distribution. Dominica, Guadeloupe, PuertoRico; widespread Antillesendemic. Notes. Adults attracted to lights. Plate 12.

Psammodius viti Chalumeau 1983c: 83 . Distribution. Les Saintes; single island endemic. Notes. Sometimes common in sand and under debris of beach of Grande Anse after heavy rain.

TRIBERHYPARINI

Rhyparus spilmani Cartwright and Chalumeau in Chalumeau 1977a: 76; Cartwright and Chalumeau 1978: 8; Chalumeau 1983a: 93; Marquet and Roguet 2003: 12; Touroult 2005: 83. Distribution. Dominica, Guadeloupe, Martinique; Lesser Antilles endemic. Notes. Collected by at light trap and by sifting plant debris. Plate 12.