Claviderma compactum, Ivanov & Scheltema, 2008

Ivanov, Dmitry L. & Scheltema, Amélie H., 2008, NMR solution structures of KAP- 1 PHD finger-bromodomain, Zootaxa 1885, pp. 1-60 : 14-15

publication ID	https://doi.org/ 10.13018/bmr11036
DOI	https://doi.org/10.5281/zenodo.5134427
persistent identifier	https://treatment.plazi.org/id/3076EC0B-FF8C-FF83-5DAB-FE6CFDFA08EB
treatment provided by	Felipe
scientific name	Claviderma compactum
status	sp. nov.

Treatment

Claviderma compactum View in CoL n. sp.

( Figures 1C View FIGURE 1 , 7 View FIGURE 7 , 8 View FIGURE 8 , 13A, B View FIGURE 13 ; Table 5)

Type material: Holotype USNM 1112622 About USNM . Paratypes: USNM; TAMU ; ZMUM Le1–172 (type locality); Le1–173 (AII-60 239); Le1–174 (AII-60 239); Le1–175 (AII-60 240); Le1–176 (AII-60 262).

Type locality: Argentine Basin , 36°12.7'S, 52°42.7'W, 2,041 m ( RV Atlantis II Cruise 60, station 264, 28.03.1971) GoogleMaps .

Material examined: Argentine Basin , 687 specimens ( Table 5) .

Diagnosis: A small Claviderma species <3 mm, shank short, fuzzy from long upright sclerites, <1/3 trunk length, appearing nearly same width as trunk, sclerites upright on an anteroventral bulge, dorsal and remaining ventral trunk sclerites adpressed, at 45º to longitudinal body axis; shank clearly demarcated from trunk but not from knob; fringing sclerites not in ring, scarcely extending beyond knob; knob with two lateral groups of long sclerites. Oral shield sclerites in 3 rows. Trunk sclerites straight, narrow, base long, tapered proximally, blade short, narrower than base, with straight tapered sides ending in sharp point.

Etymology: L. compactus, -a, -um = compact.

Description: External appearance ( Figs 1C View FIGURE 1 , 7A–C View FIGURE 7 ): General: Small, trunk smooth, sclerites raised on shank giving it fuzzy, wide appearance; holotype length 2.7 mm, anterium 0.1 mm, trunk 1.6 x 0.6 mm, shank 0.6 X 0.4 mm, knob 0.4 mm, posterium/trunk index 0.37; base of large jaws within hemocoele forming an anteroventral bulge raising sclerites upright ( Fig. 7A–C View FIGURE 7 , arrows); trunk sclerites arranged from anteroventral to dorsoposterior, diverging from ventral midline rather far posteriorly but not converging dorsally. Oral shield 0.18 mm in height.

Sclerites ( Figs 7D, G View FIGURE 7 , 8 View FIGURE 8 ): General: Blade very short above waist on anterodorsal trunk, lengthening gradually posteriorly to the shank, thickest medially on base of trunk sclerites, thick for size of sclerites. Oral shield ( Fig. 7D View FIGURE 7 , 8A View FIGURE 8 ): triangular with rounded, angular base, rounded distally, thickest proximally, 65 x 30 x> 3 µm; of nearly equal size in the 3 rows, largest dorsally. Dorsal to oral shield ( Fig. 8B View FIGURE 8 ): oval or with waist, 50 x 25 x 2 µm. Anterium ( Fig. 8C View FIGURE 8 ): asymmetrical and deeply waisted, to 100 x 25 x 3 µm. Trunk beneath anteroventral bulge ( Fig. 8D View FIGURE 8 ): 2 types, one with deep waist like anterium sclerites, the other ovoid with a short blade, to 110 x 30 x 5 µm, thickest medially on base. Ventral trunk posterior to bulge ( Fig. 8E, F View FIGURE 8 ): like second type of sclerites beneath bulge, but larger and with longer blades,>200 x 40 x 7 µm, thickest medially or distally on base. Dorsal trunk sclerites from anterior to posterior ( Fig. 8G, H View FIGURE 8 ): similar to ventral sclerites, width of base nearly the same throughout trunk, base and blade both increasing in length, with blade increasing from 1/7 to 1/4 total length, from 150 x 30 x 4 µm to 250 x 35 x 8 µm, thickest medially on base. Shank ( Fig. 8I View FIGURE 8 ): 2 types, one type with shape like trunk sclerites but with longer blade up to 1/2 total length, to 280 x 30 x 8 µm, thickest on blade; the second type shorter with a wide quadrate base, base and blade of about equal length, to 220 x 60 x 7 µm, thickest medially on blade. Fringe ( Fig. 8J View FIGURE 8 ): with quadrate base and blade about 3 times length of base, 360 x 50 x 10 µm, or needle-like, 340 x15 x 8 µm. Knob ( Fig. 8K View FIGURE 8 ): short, straight, tapered or swollen proximally, to 210 x 20 x 4 µm, or very long and hair-like, to 500 x <5 µm.

Radula and jaws: 2 examined. Jaws 520 x 200 µm ( Fig. 7E View FIGURE 7 ); radula teeth 140 µm long, serrated membrane about 90 µm long, medial; central plate symmetrical, curved, 30 x 10 µm ( Fig. 7F View FIGURE 7 ).

Distribution ( Fig. 25 View FIGURE 25 , Table 5): Known from 5 stations in the Argentine Basin at just above to within the continental rise (993–3,343 m) ( Table 5).

Remarks: Cl. compactum is distinguished from other Claviderma species by: a wide and fuzzy short shank and small size; 3 rows of oral shield sclerites almost equal in size; short, and narrow sclerite blades with straight sides tapering to a narrow proximal end. The anatomy of the relationship between jaws and hemocoele is found in Scheltema (1981).