Allanagrus Noyes & Valentine, 1989

Huber, John T. & Triapitsyn, Serguei V., 2017, Two genera of Mymaridae (Hymenoptera) new to Africa, a remarkable new species of Anaphes and new generic synonymy, ZooKeys 658, pp. 39-61 : 46

Allanagrus Noyes & Valentine, 1989

Type species.

Allanagrus magniclava Noyes & Valentine.

Allanagrus contains three described species in Australia ( Lin et al. 2007) and one in New Zealand (Noyes & Valentine 1989), but the genus also occurs in the Oriental region ( Triapitsyn 2014b). Allanagrus species have the clava 3-segmented (or apparently so in some doubtful cases) and tarsi 4-segmented, both features shared by at least one species of 18 other genera: Allarescon Noyes & Valentine, Anaphes , Anneckia Subba Rao, Eustochus Haliday, Kompsomymar Lin & Huber, Krokella Huber, Neostethynium Ogloblin, Nesomymar Valentine, Nesopatasson Valentine, Notomymar Doutt & Yoshimoto, Paracmotemnus Noyes & Valentine, Paranaphoidea , Parastethynium Lin & Huber, Platystethynium Ogloblin, Polynemoidea Girault, Pseudanaphes Noyes & Valentine, Pseudocleruchus Donev & Huber, and Stethynium Enock. These genera are not necessarily related, however. Three genera, Nesomymar , Nesopatasson and Notomymar , are only known from wingless specimens whose relationships are uncertain-these genera may be represented also by winged species described in other genera. The genera most similar to Allanagrus appear to be Anneckia , Parastethynium (both in Australasian region) and Stethynium (almost worldwide) based on: face with subantennal grooves, and frenum weakly sclerotized and apparently divided medially by a longitudinal groove. The strongly oblique suture of the compact clava in Stethynium removes this genus from further consideration here. The new species described below differs from Anneckia in that the campaniform sensilla of the dorsellum abut the anterior margin, as in one of the two species of Parastethynium (well separated from anterior margin in Anneckia and in the type species of Parastethynium ) and the mandible (female) is well developed and probably has 2 distinct ventral teeth (not clearly visible in holotype of new species) and a dorsal, serrated edge somewhat as in Parastethynium (1 tooth in the reduced mandible of Anneckia ). It differs from Parastethynium in that the fore wing is fairly narrow with rounded apex, and the hind wing is narrow (fore wing wide and apically truncate and hind wing quite wide in Parastethynium ) and eye is much less setose (eye with many setae in Parastethynium ). On balance of features we tentatively place the new species in Allanagrus . A detailed study of all these genera is needed to clarify their relationships and determine if some should be synonymized under others or, conversely, more should be proposed.