Scydmaenus (Nepaloscydmaenus) kiautunensis Franz, 1985

Jałoszyński, Paweł, 2021, Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4952 (2), pp. 354-368 : 364

Scydmaenus (Nepaloscydmaenus) kiautunensis Franz

Scydmaenus (Scydmaenus) kiautunensis Franz, 1985: 117 .

( Figs 31–36 )

Type material. Holotype ( CHINA): ♂, three labels ( Fig. 36 ): “KUATUN,FUKIEN / China 8.5.46 / (TSCHUNG SEN.)” [white, printed and handwritten], “ Scydmaenus / kiautunensis m. / det. H.Franz ” [white, handwritten and printed], “ Holotypus ” [red, handwritten] ( NHMW).

Revised diagnosis. Male. Legs and metaventrite unmodified, but elytral apex with large circular pit with dense setae behind it; head virtually impunctate, pronotum with fine but distinct and very dense punctures, elytra with yet larger but sparser, distinct punctures; head about as wide as long, with small eyes; tempora in dorsal view slightly more than 2.5 × as long as eye; pronotum slightly longer than wide, lacking antebasal pits; aedeagus in dorsal view gradually tapering from base to apex but with slightly concave sides in more than median third, apical margin with distinct subtriangular emargination separating rounded lateral lobes, in lateral view distal region of median lobe distinctly recurved with long and flat apical projection bent dorsad; sides of ostium lacking rows of setae, but lateral margins of median lobe in subapical region bear several long setae; endophallus with long tubular projection rapidly curved or bent several times.

Redescription. Body of male ( Fig. 25 ) elongate and strongly convex, uniformly brown, covered with light brown setae; BL 1.89 mm.

Head in dorsal view broadest at eyes, about as wide as long, HL 0.36 mm, HW 0.35 mm; vertex and frons confluent, evenly convex; supraantennal tubercles feebly marked; tempora in dorsal view slightly more than 2.5 × as long as eye; eye strongly convex, in lateral view weakly oval. Frons and vertex virtually impunctate; setae relatively long and dense, suberect. Antenna slender, AnL 1.00 mm; antennomeres 1–8 each strongly elongate, 9 about as long as broad, 10 indistinctly elongate, 11 indistinctly shorter than 9–10 combined, slightly more than twice as long as broad.

Pronotum in dorsal view broadest near anterior third; PL 0.50 mm, PW 0.45 mm; anterior margin nearly straight, sides in anterior half rounded, in posterior half nearly straight and strongly converging posterad; posterior margin nearly straight; base lacking pits Pronotal disc with fine and shallow but sharply marked punctures separated by spaces narrower than their diameters; setae short, sparse and erect.

Elytra together oval, broadest in front of middle (based on shape of one elytron), EL 1.03 mm, EW 0.70 mm, EI 1.46. Humeral calli barely discernible; basal impressions indistinct, anterior portion of suture not elevated and not demarcated by grooves; elytral apex with large circular pit closer to sutural than to lateral margin and separated from apical margin by slightly less than its diameter. Elytra covered with punctures more distinct, larger and deeper than those on pronotum and separated by spaces narrower than their diameters; setae slightly thicker than those on head and pronotum and distinctly longer than those on pronotum, moderately dense and long, suberect, not forming waving pattern, those behind apical pit forming dense patch directed mesad. Hind wings in the holotype seem shortened, but the elytra have been disarticulated during previous preparations and wings might have been damaged then.

Legs long and conspicuously slender, unmodified, including slender, not broadened protarsi.

Metaventrite unmodified.

Aedeagus ( Figs 32–35 ) moderately elongate, AeL 0.45 mm; median lobe in dorsal view gradually tapering from base to apex but with slightly concave sides in more than median third, apical margin with distinct subtriangular emargination separating rounded lateral lobes, in lateral view basal half of median lobe much broader than distal half, distal region bulging dorsad in subapical area and distinctly recurved with long and flat apical projection bent dorsad; sides of ostium lacking rows of setae, but lateral margins of median lobe in subapical region with several long setae; endophallus with long tubular projection rapidly curved or bent several times.

Female. Unknown.

Distribution. Fujian.

Remarks. This species was originally placed in Scydmaenus s. str. However, Franz (1973) established a subgenus Nepaloscydmaenus , which differs from other subgenera in elongate and slender antennomeres 7 and 8, antennomere 8 very narrow, trimerous and sharply demarcated club, and all antennomeres symmetrical (orig.: “Von den bekannten Subgenera der Gattung Scydmaenus durch längliches und schmales 7. und 8. Fühlerglied verschieden, das 8. Fühlerglied sehr dünn, die grosse 3-gliederige Keule von ihm scharf abgesetzt. Alle Fühlerglieder symmetrisch gebaut”). Additionally, S. ghokarnae Franz, 1973 , the type species of Nepaloscydmaenus , has the pronotum lacking antebasal pits, the metanepisterna demarcated from the metaventrite, and the protarsomeres in male not broadened. The type material of S. ghokarnae (NHMW) was compared with the holotype of S. kiautunensis , and both species clearly share the same characters diagnostic for Nepaloscydmaenus . Consequently, S. kiautunensis is transferred from Scydmaenus s. str. to Nepaloscydmaenus . The holotype of S. kiautunensis is incomplete, the left elytron is missing; the width of elytra was interpolated from measurements of the right elytron before remounting, resting in natural position.

Figures

FIGURES 31–36. Scydmaenus (Nepaloscydmaenus) kiautunensis Franz, holotype male. Dorsal habitus (31); aedeagus in dorsal (32, 34) and lateral (33, 35) views; original labels (36).

FIGURES 25–30. Scydmaenus (Scydmaenus) szechuanensis Franz, holotype male. Dorsal habitus (25); aedeagus in dorsal (26, 28) and lateral (27, 29) views; original labels (30).