Myrmeleotettix I. Bolívar, 1914

Sergeev, M. G., Storozhenko, S. Yu. & Benediktov, A. A., 2019, An annotated check-list of Orthoptera of Tuva and adjacent regions. Part 2. Suboder Caelifera. Tridactylidae, Tetrigidae, Acrididae: Melanoplinae, Calliptaminae, and Gomphocerinae (except Gomphocerini), Far Eastern Entomologist 389, pp. 7-44 : 29-32

Genus Myrmeleotettix I. Bolívar, 1914

35(57). Myrmeleotettix palpalis (Zubovsky, 1900)

Omocestus (Myrmeleotettix) palpalis: Berezhkov, 1951: 18 .

Myrmeleotettix palpalis: Berezhkov, 1956: 98 ; Ivanova, 1967: 131; Sergeev, 1986: 205; Sergeev et al., 1995: 96– 98; Benediktov, 1997: 117; Pavlov, 2004: 65; Tishechkin & Bukhvalova, 2009: 27.

Myrmeleotettix pallasi (sic!): Kazakova & Sergeev, 1993: 73.

MATERIAL. Tuva: W Sayan Mts., Alash Plateau, Ak-Sug River, 51°24ʹN, 90°27ʹE, 1079–1085

m, southern slope, stony semi-desert, 13.VIII 2016, 3 ♂, 16 ♀ ( MS); W Sayan Mts. , Alash Plateau, AkSug River, 51°31ʹN, 90°12ʹE, 1275 m, southern slope, steppe, 14.VIII 2016, 4 ♀ ( MS); W Sayan Mts .,

Alash Plateau, Ak-Sug River, 51°30ʹN, 90°12ʹE, 1250–1252 m, lower terrace and upper flood-plain,

meadows, pebbles, 14.VIII 2016, 1 ♂, 19 ♀ ( MS); Turan-Uyuk Intermountain Basin , Begreda River ,

51°59ʹN, 94°18ʹE, 830–845 m, southern slope and piedmont plain, semi-deserts, 12.VIII 2018, 4 ♀ ( MS) ;

Chaa-Khol River , 51°34′N, 92°23′E, 570–600 m, dry steppes, 14–18.VII 1962, 24 ♂, 49 ♀, 9 larvae ( IS) ;

14 km SW Shagonar City, Shagonar River , 51°27ʹN, 92°45ʹE, 583 m, lower terrace, meadow between bushes, 17–18.VI 2017, 4 ♂, 1 ♀, 7 larvae ( MS) ; 14 km SW Shagonar City, Shagonar River , 51°28ʹN ,

92°44ʹE, 615–629 m, slopes and piedmont plain, semi-deserts, 21, 23.VI 2017, 54 ♂, 8 ♀, 46 larvae

( MS); 14 km SW Shagonar City, Shagonar River , 51°28ʹN, 92°45ʹE, 585–600 m, semi-desert and dry steppe, 25–26.VI 2017, 26 ♂, 5 ♀, 15 larvae ( MS) ; 32 km SW Kyzyl City, Elegest River , 51°29ʹN ,

94°10ʹE, steppe, 22.VII 2014, 6 ♂, 10 ♀ ( SS); Ulug-Khem Intermountain Basin , 25 km SW Kyzyl City ,

Ulug River , 51°35′N, 94°09′E, 650–700 m , piedmont plain and upper terraces, dry steppes and semidesert, 6.VII 1978, 16 ♂, 9 ♀, 16 larvae ( MS); 25 km SW Kyzyl City, Ulug River , 51°36′N, 94°09′E ,

600–605 m, upper terrace, dry steppe, 7.VIII 1978, 3 ♂, 2 ♀, 6 larvae ( MS); 25 km SW Kyzyl City, Ulug

River, 51°36′N, 94°09′E, 600–605 m, lower terrace, steppe, 7.VIII 1978, 12 ♂, 11 ♀, 3 larvae ( MS) ;

Ulug-Khem Intermountain Basin, 25 km SW Kyzyl City, Ulug River, 51°35ʹN, 94°09ʹE, 630–830 m,

mountain slopes and piedmont plain, semi-deserts Nanophyton grubovi , 19.VII 1995, 21 ♂, 11 ♀ ( MS) ;

Ulug-Khem Intermountain Basin , 25 km SW Kyzyl City, Ulug River, 51°35ʹN, 94°08ʹE, 608 m, upper terrace, semi-deserts Nanophyton grubovi , 20.VII 1995, 3 ♂, 1 ♀ ( MS); Ulug-Khem Intermountain

Basin, 25 km SW Kyzyl City, Ulug River , 51°36ʹN, 94°09ʹE, 600–605 m, lower terrace and upper floodplain, meadows, 20.VII 1995, 1 ♂ ( MS); Ulug-Khem Intermountain Basin , 25 km SW Kyzyl City, Ulug

River, 51°34ʹN, 94°10ʹE, 800–900 m, southern slope and piedmont plain, semi-deserts Nanophyton grubovi , 22.VII 2003, 24 ♂, 22 ♀, 2 larvae ( MS); Ulug-Khem Intermountain Basin , 8 km N Cheder Lake ,

51°32ʹN, 95°21ʹE, 750–760 m, steppes, 12.VIII 1962, 3 ♀ ( IS); SE Ulug-Khem Intermountain Basin ,

Uzun-Kharagan River , near Balgazyn settlement, 50°57ʹN, 95°16ʹE, 940 m, southeastern slope, stony steppe, 8.VII 1978, 10 ♂, 13 ♀, 12 larvae ( MS); SE Ulug-Khem Intermountain Basin, Uzun-Kharagan

River, near Balgazyn settlement, 50°57ʹN, 95°16ʹE, 940 m, upper terrace, meadow with Caragana bushes, 8.VII 1978, 19 ♂, 24 ♀ ( MS); SE Ulug-Khem Intermountain Basin, Shuurmak River , 25 km S

Balgazyn settlement, near Kuran settlement, 50°47ʹN, 95°17ʹE, 1030–1050 m, southern slope, steppe and meadow, 10.VII 1978, 18 ♂, 22 ♀, 7 larvae ( MS); W Tannu-Ola Mts. , 5 km N Khandagajty settlement, 50°48ʹN, 92°05ʹE, 1300–1350 m, piedmont plain, stony steppes with Caragana , 16.VIII 1985 ,

22 ♂, 32 ♀ ( MS); W Tannu-Ola Mts. , 5 km N Khandagajty settlement, Mugur River, 50°46ʹN, 92°04ʹE ,

1170 m, marsh, wet meadow with bushes, 17.VIII 1985, ♀ ( MS); W Tannu-Ola Mts. , 5 km N

Khandagajty settlement, 50°47ʹN, 92°02ʹE, 1225 m, piedmont plain, stony steppes with Caragana ,

18.VIII 1985, 7 ♂, 4 ♀ ( MS); W Tannu-Ola Mts. , 9 km N Khandagajty settlement, 50°51ʹN, 92°04ʹE ,

1700–1900 m, mountain steppes, 19.VIII 1985, 3 ♂, 8 ♀ ( MS); W Tannu-Ola Mts. , 9 km N Khandagajty settlement, 50°50ʹN, 92°05ʹE , 1637 m, mountain steppes, 19.VIII 1985, 5 ♂, 7 ♀ ( MS); W Tannu-Ola

Mts., NW Khandagajty settlement, near Solchur settlement, 50°46ʹN, 92°01ʹE, 1150–1200 m,

agricultural fields including abandoned and plots along canal and roads, 21.VIII 1978, 6♂, 12 ♀ ( MS) ;

W Tannu-Ola Mts. , 5 km N Khandagajty settlement, 50°47ʹN, 92°03ʹE, 1193 m, piedmont plain, lower terrace of stream, meadow, 21.VIII 1985, 4 ♂, 6 ♀ ( MS); W Tannu-Ola Mts., 5 km N Khandagajty settlement, 50°47ʹN, 92°05ʹE, 1200–1250 m, piedmont plain, lower part, stony steppes with Caragana and dry meadow, 21.VIII 1985, 16 ♂, 31 ♀ ( MS); W Tannu-Ola Mts., 5 km N Khandagajty settlement ,

50°48ʹN, 92°04ʹE, 1336 m, foothills on piedmont plain, stony steppe, 21.VIII 1985, 2 ♂, 3 ♀ ( MS); W

Tannu-Ola Mts., 5 km N Khandagajty settlement, 50°50ʹN, 92°05ʹE, 1500-1550 m, piedmont plain,

upper part, steppe, 21.VIII 1985, 7 ♂, 11 ♀ ( MS); W Tannu-Ola Mts. , 4 km E Khandagajty settlement ,

50°45ʹN, 92°09ʹE, 1150–1200 m, southern slope and piedmont plain, stony semi-desert with Nanophyton grubovii , 24.VIII 1985, 4 ♂, 11 ♀ ( MS); W Tannu-Ola Mts. , near Khandagajty settlement, 50°44′N ,

92°09′E, 1140 m, northern slope, stony semi-desert, 24.VIII 1985, 1 ♂, 2 ♀ ( MS); W Tannu-Ola Mts. , near

Khandagajty settlement, 50°43′N, 92°04′E, 1170–1200 m, piedmont plain, desert with Nanophyton grubovii , 24.VIII 1985, 1 ♂, 3 ♀ ( MS); W Tannu-Ola Mts. , near Khandagajty settlement, 50°43′N ,

92°03′E, 1200–1250 m, piedmont plain and northern slope, semi-desert with Caragana bushes, 24.VIII

1985, 1 ♀ ( MS); W Tannu-Ola Mts. , 5 km W Torgalyg settlement, Khandybai Mt., mountain steppe ,

23.VI 1962, 1 larva ( IS); E Tannu-Ola Mts. , Shivelig River, 50°45ʹN, 94°34ʹE, 1330–1340 m, flood-plain and terraces, steppe and short meadow between shrubs and Larix trees , 28.VI 1978, 42 ♂, 13 ♀, 366

larvae ( MS); E Tannu-Ola Mts. , Shivelig River, 50°45ʹN, 94°34ʹE, 1330–1340 m, lower terrace, steppe and short meadow between shrubs and Larix trees, 17.VII 1978, 3 ♂, 2 ♀ ( MS) ; the same locality and habitat, 12.VIII 1978, 1 ♂, 7 ♀ ( MS) ; the same locality, upper terrace, steppe, 12.VIII 1978, 31 ♂, 96 ♀

( MS); the same locality, upper flood-plain, meadow, 12.VIII 1978, 1 ♂ ( MS); E Tannu-Ola Mts. , Shivelig

River, 50°45ʹN, 94°34ʹE, 1330–1340 m, upper terrace, steppe and short meadow with Caragana , 17.VII

1978, 57 ♂, 148 ♀, 29 larvae ( MS); the same locality, southern slope and upper terrace, stony steppes ,

17.VII 1978, 54 ♂, 133 ♀, 40 larvae ( MS); E Tannu-Ola Mts. , Shuurmak River, 5 km N Shuurmak settlement, 50°41ʹN, 95°19ʹE, 1173 m, local southern slope and piedmont plain, steppe , 10.VII 1978, 14 ♂,

13 ♀ ( MS); E Tannu-Ola Mts. , Shuurmak River, 5 km N Shuurmak settlement, 50°42ʹN, 95°20ʹE, 1106

m, southern slope, mountain steppe, 11.VII 1978, 54 ♂, 76 ♀, 1 larva ( MS); E Tannu-Ola Mts .,

Shuurmak River, 5 km N Shuurmak settlement, 50°42ʹN, 95°20ʹE, 1086 m, lower terrace, meadow,

11.VII 1978, 1 ♂ ( MS); E Tannu-Ola Mts. , Shuurmak Pass, 50°37ʹN, 95°11ʹE, 1495 m, southern slope and upper terrace, mountain steppe , 12.VII 1978, 116 ♂, 104 ♀ ( MS); N Uvs-Nuur Intermountain

Basin , Iribitej River, 50°44′N, 93°08′E, 938–973 m, piedmont plain, stony semi-deserts with

Nanophyton grubovii , 24.VI 1978, 27 larvae ( IS, MS); N Uvs-Nuur Intermountain Basin, Iribitej River ,

50°45′N, 93°09′E, 1030–1040 m, terraces, dry meadows and steppes, 29.VII 1978, 6 ♂, 7 ♀ ( MS); N

Uvs-Nuur Intermountain Basin , Iribitej River, 50°44′N, 93°08′E, 984 m, upper terrace, stony semi-desert ,

29.VII 1978, 8 ♀ ( MS); N Uvs-Nuur Intermountain Basin, Iribitej River , 50°44′N, 93°08′E, 938 m ,

lower terrace and flood-plains, steppe, 30.VII 1978, 8 ♂, 25 ♀, 2 larvae ( MS); N Uvs-Nuur

Intermountain Basin , Iribitej River, 50°44′N, 93°08′E, 938 m, upper terrace and piedmont plain, semidesert with Nanophyton grubovii , 30.VII 1978, 12 ♂, 26 ♀ ( MS); N Uvs-Nuur Intermountain Basin ,

Iribitej River , 50°44′N, 93°08′E, 973 m, piedmont plain, upper part, semi-desert with Nanophyton grubovii , 30.VII 1978, 5 ♂, 3 ♀ ( MS); N Uvs-Nuur Intermountain Basin, 50 km E Amdaygyn-Khol ,

semi-desert with Nanophyton grubovii , 26.VII 1962, 6 ♂, 4 ♀ ( IS); N Uvs-Nuur Intermountain Basin ,

40 km E Amdaygyn-Khol , semi-desert with Nanophyton grubovii , 26.VII 1962, 2 ♀ ( IS); N Uvs-Nuur

Intermountain Basin, Amdaygyn-Khol, 50°42ʹN, 93°16ʹE, 783 m, meadow, 27.VII 1962, several specimens ( IS); NE Uvs-Nuur Intermountain Basin, Khol-Oozhu River, 50°43ʹN, 94°17ʹE, 931 m, piedmont plain, lower part, semi-desert with Nanophyton grubovii , 30.VI 1978, some specimens observed ( MS);

NE Uvs-Nuur Intermountain Basin, Khol-Oozhu River, 50°43ʹN, 94°17ʹE, 930 m, upper flood-plain,

meadow, 16.VII 1978, 1 ♂ ( MS); NE Uvs-Nuur Intermountain Basin, Khol-Oozhu River , 50°43ʹN ,

94°17ʹE, 930 m, piedmont plain, lower part, semi-desert with Nanophyton grubovii , 16.VII 1978, 8 ♂, 3

♀, 8 larvae ( MS); NE Uvs-Nuur Intermountain Basin, Shivelig River , 50°39ʹN, 94°28ʹE, 990–995 m ,

flood-plain and terraces, dry steppes with Caragana , 28.VI 1978, 32 larvae ( MS); the same locality and habitats, 18.VII 1978, 62 ♂, 42 ♀, 46 larvae ( MS); the same locality and habitats, 10.VIII 1978, 13 ♂ , 21

♀, 1 larva ( MS); NE Uvs-Nuur Intermountain Basin, Shivelig River , 50°43ʹN, 94°33ʹE, 1215 m, lower terrace, dry steppe, 15.VII 1978 , 8 ♂, 9 ♀, 6 larvae ( MS); the same locality and habitats, 11.VIII 1978 , 3

♂, 11 ♀ ( MS); the same locality, 1200–1300 m, lower terrace, meadow 15.VII 1978 , 1 ♀, 1 larva ( MS); NE

Uvs-Nuur Intermountain Basin , Shivelig River, 50°43ʹN, 94°33ʹE, 1200–1300 m, southern slopes and piedmont plain, steppes with Caragana , 24.VII 2003, 5 ♂, 3 ♀, 6 larvae ( MS); NE Uvs-Nuur

Intermountain Basin, Shivelig River, 50°45ʹN, 94°34ʹE, 1280–1301 m, terraces, dry meadows, 8.VII

2017, some specimens observed ( MS); NE Uvs-Nuur Intermountain Basin, Shivelig River, 50°43ʹN,

94°33ʹE, 1259 m, piedmont plain, dry steppe with Caragana , 8.VII 2017, 22 ♂, 14 ♀, 3 larvae ( MS); NE

Uvs-Nuur Intermountain Basin, Shivelig River, 50°43ʹN – 50°45ʹN, 94°33ʹE – 94°34ʹE, 1200–1650 m,

mountain slopes, piedmont plains, terraces, dry meadows and steppes, often with bushes and stones,

24.VI–12.VIII 1978,> 200 specimens, including larvae ( MS, T. Myagkaya, A. Lee); Uvs-Nuur Intermountain Basin, Tes River , 50°33ʹN, 94°31ʹE, 932–934 m, upper flood-plain and terraces, dry meadows and steppes , 27.VI 1978, 31 ♂, 1 ♀, 178 larvae ( MS); Uvs-Nuur Intermountain Basin, Tes River , 50°32ʹN ,

94°31ʹE, 930–932 m, upper flood-plain, meadows, 13.VII 1978, 17 ♂, 12 ♀, 9 larvae ( MS); Uvs-Nuur

Intermountain Basin, Tes River, 50°33ʹN, 94°31ʹE, 932–934 m, terraces, steppes, 14.VII 1978, 45 ♂, 45

♀, 38 larvae ( MS); Uvs-Nuur Intermountain Basin , Tes River, 50°32ʹN, 94°31ʹE, 930–932 m, upper flood-plain and terraces, 11.VIII 1978 , 10 ♂, 23 ♀ ( MS); Dyttyg-Khem River , 12 km SW Samagaltai settlement, 50°38ʹ N, 95°19ʹ E, 17.VII 2014 , 3 ♂, 5 ♀ ( SS); Dyttyg-Khem River , 13 km SW Samagaltai settlement, 9.VII 2013 , 1 ♀ (M. Prostshalykin, V. Loktionov ); Uvs-Nuur Intermountain Basin, DusKhol, 50°22ʹN, 94°52ʹE, 994–995 m, semi-desert with Nanophyton grubovii , 24–26.VII 1962 , 7 ♂, 11 ♀

( IS); Uvs-Nuur Intermountain Basin, Dus-Khol, 50°23ʹN, 94°52ʹE, solonchak, 13.VII 2014, 3 ♂, 3 ♀

( SS); Uvs-Nuur Intermountain Basin, 7 km S Erzin settlement, Tes River, 50°11ʹN, 95°12ʹE, 1097 m,

lower terrace and upper flood-plains, dry meadows and steppes, 22.VII 1978, 1 ♂, 3 ♀ ( MS); Uvs-Nuur

Intermountain Basin, 7 km S Erzin settlement, Tes River, 50°11ʹN, 95°12ʹE, 1097 m, lower terrace,

steppes, 23.VII 1978, 7 ♂, 17 ♀, 2 larvae ( MS); 7 km SW Erzin settlement, Tes River , 50°12ʹN ,

95°08ʹE, steppe, 13.VII 2014, 5 ♂, 3 ♀ ( SS); Uvs-Nuur Intermountain Basin , 10 km S Erzin settlement ,

Tsuger-Els, 50°10ʹN, 95°11ʹE, 1154 m, sandy semi-desert, 23.VII 1978, 1 ♂, 5 ♀, 1 larva ( MS); UvsNuur Intermountain Basin, E Shara-Nuur, Yamalyg farwell rocks, 50°14ʹN, 94°45ʹE, 1150 m, piedmont plain, dry steppe, 6.VII 2017, some specimens observed ( MS); Uvs-Nuur Intermountain Basin, near Shara

Lake, 50°13ʹN, 94°32ʹE, 902–904 m, plain and upper terrace, sands, semi-desert and grassland with

Achnatherum , 6.VII 2017, 1 ♂ ( MS); Uvs-Nuur Intermountain Basin , near Shara Lake, 50°14ʹN ,

94°31ʹE, 894 m, flood-plains of lake and small stream, short meadows with halophytes, 6.VII 2017, 1 ♂

( MS); 25 km SW Erzin settlement, Tes River , 50°05ʹN, 95°21ʹE, 14–15.VII 2014, 1 ♀ ( SS); 25 km SSW

Erzin settlement, Tore Lake, 50°02ʹN, 95°03ʹE, 1150 m, sands, grasses and Caragana , 11–12.VII 2014,

12 ♂, 14 ♀ ( SS); Tore Lake , 50°06ʹN, 95°06ʹE, 1100 m, coast, steppe, 12.VII 2014 , 6 ♂, 6 ♀ ( SS); UvsNuur Intermountain Basin, S Tore Lake , 50°01′N, 95°04′E, 1154-1166 m, 07.VII 2017, sand semi-desert and upper terraces with birch forest , 2 ♂ ( MS); Uvs-Nuur Intermountain Basin, S Tore Lake , 50°05′N ,

95°09′E, 1174 m, sandy semi-desert with Caragana bungei , 7.VII 2017, 6 ♂, 1 ♀, 1 larva ( MS); 31 km

NEE Erzin settlement, Erzin River, 50°21ʹN, 95°34ʹE, 1300 m, northern slope, balka with Caragana ,

18.VII 2014, 1 ♂, 2 ♀ ( SS); 31 km NEE Erzin settlement, Erzin River , 50°21ʹN, 95°34ʹE, 1100 m, floodplain, meadow near cliff , 18.VII 2014, 2 ♂, 1 ♀ ( SS); 34 km NEE Erzin settlement, Erzin River , 50°22ʹN ,

95°30ʹE, 1100 m, larch forest, openings, 16.VII 2014, 1 ♂, 1 ♀ ( SS); 25 km NEE Erzin settlement, Belyj

Medved Mt., 50°21ʹN, 95°27ʹE, steppe, 16.VII 2014, 2♂, 3 ♀ ( SS). Krasnoyarsk Region: southern part,

West Sayan Mts. , Us Intermountain Basin, near abandoned Idzhim settlement, 52°16′N, 93°07′E , 680–

700 m, terraces and southern slope, steppes, 22–23.VII 1995, 1 ♂ ( MS) .

DISTRIBUTION. Tuva: WSW, WSE, KHE, UKH, KKH, WTO, ETO, UVS, SAN. – S

Siberia (from the Altay Mts. to Dauria), Amur Region; E Kazakhstan, Mongolia, China.

ECOLOGY. One of the most common and abundant grasshoppers in the dry steppes and semi-deserts.