Cyana dohertyi (Elwes, 1890)

Volynkin, Anton V., Černý, Karel, Saldaitis, Aidas & Ivanova, Maria S., 2019, On the correct identification of ‘ Cyana dohertyi (Elwes, 1890) ’ from northern Indochina and China, with descriptions of two new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae), Zootaxa 4658 (1), pp. 155-167 : 156

The C. dohertyi species-group

Diagnosis. The species-group is characterized by having yellow or yellowish-red transverse lines, by the presence of a ventral subbasal cluster of spine-like cornuti and a largest diverticulum with (or without) a cluster of spinules in the male vesica, and large bulbous or elliptical appendix bursae with a narrow and heavily sclerotized proximal section in the female genitalia ( Volynkin et al. 2017).

Morphology. Adults. Medium-sized moths with forewing length 14–21 mm in males and 17.5– 25 mm in females. The sexual dimorphism well expressed: besides presence of forewing androconial lobes on the underside of forewing, males have smaller size, narrower forewing, presence of black stroke on R 2 vein, and usually darker hindwing. Head white. Antennae ciliate in both sexes. Thorax white, with broad orange bands on tegulae and patagia margins, and orange transverse bands on medial and posterior parts. Abdomen whitish, in some species with pinkish suffusion. Forewing ground color white. Pattern consists of transverse lines and black dots, in some species ( C. divakara and C. gazella ) transverse lines fused into shades of various shape. Transverse lines orange, wavy. In apical part of cell there are three large black discal spots. Cilia white. Hindwing from white to pale red- dish; discal spot semilunar, poorly visible; cilia as ground color. Male genitalia. Uncus short, slender, apically narrowed, basally connected with scaphium. Anal tube broad, scaphium narrow, weakly sclerotized, subscaphium broad, setose. Tegumen of moderate length, narrow; vinculum short, broad, U-like; juxta trapezoidal, with weaker sclerotized medial part. Valva lobe-like, with strongly curved dorsal margin, distally narrowed; sacculus broad, moderately sclerotized; distal saccular process long, robust, with spine-like tip, distally curved and well separated from distal part of valva. Ventral plate of costa heavily sclerotized, with short crest. Aedeagus long and narrow, carina with short triangular extension dorsally and field of scobination ventrally. Vesica membranous, consists of largest diverticulum (with cluster of spinules or without it), one subbasal diverticulum bearing a cluster of spinules, and 2–4 smaller distal diverticula; 1 st distal diverticulum usually bears cluster of spinules or granulation; one more, lateral diverticulum at base of the largest diverticulum; basal plate of vesica ejaculatorius long and narrow, weakly sclerotized; vesica ejaculatorius ventrally projected. Female genitalia. Ovipositor short and broad. Papillae anales rectangular with rounded corners; apophyses posteriores thin, about three times longer than apophyses anteriores; apophyses anteriores broadly triangular; ostium bursae broad, antrum funnel-like, membranous; ductus bursae of moderate size, weakly rugose; corpus bursae elongated, its posterior section strongly sclerotized, rugose; anterior section membranous, slightly rugose, with weak granulation and small elliptical or round signum. Appendix bursae large, arising leftward from posterior end of corpus bursae, distally globular and membranous, basally narrow and strongly rugose sclerotized.