Ceroptera moroccana, Papp, LÁsZLÓ, 2014

Papp, LÁsZLÓ, 2014, A Review Of The Old World Species Of Ceroptera Macquart, 1835 (Diptera, Sphaeroceridae), Acta Zoologica Academiae Scientiarum Hungaricae 60 (2), pp. 109-155 : 141-143

Ceroptera moroccana sp. n.

( Figs 78–83 )

Holotypemale ( HNHM, abdomenprepared, inapolyethylenemicrovialwithglycerol): MOROCCO, Foum Zquid, 580 m, N30°04.491’, W 06°21.616’ – homokbuckás [sand dunes] Scarabeusról [onScarabaeus], leg. Szappanos.

Paratypes ( HNHM): 2 males 3 females: samelabels .

Measurementsinmm: bodylength 2.52 (holotype), 2.07–2.87 (paratypes), wing length 2.59 (holotype), 2.07–3.25 (paratypes), wingwidth 1.00, 0.89–1.16.

Body dark greyish brown, covered by very short and dense dark microtrichia. Frons betweenorbitaliaandfrontaltrianglereddish. Parafacialiaandgenaarealsogreyishredin somefemales. Pedicelandbasal 2/3 offirstflagellomeregreyishred. Antennadorsally 0.23 mm, pedicel 0.09 mm, first flagellomere 0.11 mm. First flagellomere with 0.035–0.04 mm longslightlycurvedgreyhairs. Widthoffirstflagellomere 0.14 mm, i.e. broader (deeper) than long. Longitudinal axis of eye 0.33 mm, width of eye 0.31 mm, width of gena 0.19 mm. Facialplateshort, mouthedgestronglyprotruding, antennaelieindeepimpressions, arathernarrow, almostsharpcarinapresent. Eyesnotwidelydeparted. Either poc or occi strong. Only 4 pairsofnotshortbutalmostcolourlessinterfrontalsetae. Aristaalmost bare(!). Dorsalapicalsetaofpedicel 0.10 mm.

Anepisternumwith 3 distinctverticalfossae. Only 1 dorsocentralpair, prescutellar acrostichalsnotenlarged, 6–8 unarrangedacrostichalrow. 1 longkatepisternalsetaof 0.24 mm and other 2 short anterior katepisternals of c. 0.09 mm.

Femoraandtibiaedarkbrownwithreddishapicesandknees, tarsireddishyellow. Midtibiawithoutposterodorsaloranteroventralseta, 1 stronganterodorsalat 42/54 and adorsalsetaat 47/54. Apicalventralspurofhindtibianotparticularlystrong, 0.09 mmonly.

Wingmembranelightyellowish, veinsyellow. Secondcostalsection 0.31 mm, third section 0.48 mm, index 2.76. Vein R 4+5 slightly upcurved and ends 0.25 mm before wing apex. Costal vein overruns apex of R 4+5 considerably (by 0.07–0.09 mm). Rs 0.26 mm, inter- crossvein section of M 0.38 mm, terminal section 1.21 mm, M-M crossvein 0.19 mm. Both posterior edges of discal cell 90°, lower edge continued on a section of 0.66 mm to wing margin, veindiscerniblealmosttoit. Alulabroad, 0.10–0.21 mm.

1+2

Abdominalscleritesnotreduced. Malesyntergitewell- sclerotised, onlyanterior medialpartofT1 portiondesclerotised, medialpartofT2 portion (exceptforaposterior view. Scales: 0.2 mm for Figs 78–81, 83 , 0.1 mm for Fig, 82.

stripe) unpigmentedonabroadU-shapedarea. Setosityofthetergitesaswellasintersegmentalmembranesparselysetose. Syntergite 0.35 mmlong, 0.60 mmbroad (i.e. thatofthe abdomen). Anterior ¾ ofsternite 2 unpigmented. Measurementsofpreabdominalsclerites (1 unit = 0.0125 mm): tergite 3 17 × 43, T4 17 × 36, T5 15 × 34, S2 17 × 32, S3 20 × 40, S4 23 × 40. Sternite 5 isthemoastcharacteristicpartofmalegenitallia: thoughcomparativelylong, muchbroaderthanlong ( Fig. 78 ), posterioredgeslightlydorsallycurved, caudal ¼ with somestrongsetaeandnumerousthickbluntblackthorns. AlargemedialpartofS5 bare. Synsternitedefinitelysmall. Sternite 6 portionshortbutcontinuedinlargerightsidesclerites. Sternite 7 part with dorsally curved right part and with some 5 setae (2 of them long), sternite 8 portionatlongestonly 0.16 mm.

Epandriumnotsmall, dorsally 0.13 mmlong. Epandriumwithsparsemediumlengthsetae. Hypandriumsmallwithalongtriangularmedialpart (only 0.04 mmlong). Subepandrialsclerite ( Fig. 83 ) notbroad. Cercusroundish, ventrallybroadlyroundedand lesslong. Cercifusedsagittallywitheachotherandwithsubepandrialsclerite, withalong lateroventralsetalpairof 0.10 mm. Surstylus ( Figs 79–80 ) deeplybilobed: connectionis a 0.05–0.06 mm broad horizontal part, which isd only 0.03 mm high in profile. Since the lobesarenotparallel, depictedon 2 figures. Anteriorlobebroadeningdistally, broadly roundedwithnumerousmedium-longsetaeandwithsparsemicrotrichia. Apicalpartdirectedobliquelymedially. Posteriorlobebroadestinasubcaudal- sublateralview, basal 1/3 widened, apicalpartnarrowedwithananteriorlydirected, almostsharpapex. Posterior lobealmostbare. Genitaliasmall, e.g. phallapodemeonly 0.20 mm, distalpartslightly proclinate. Basiphallus ( Fig. 81 ) longwithadistinctepiphallus. Distiphallusnarrow, short only 0.20 mmlong, notstronglysclerotisedwithmembranousapex ( Fig. 81 ). Postgonite ( Fig. 82 ) rather large with broad rounded apex.

Femalepreabdominalscleritessmallerthanthoseofthemale. Syntergite 0.35 mm long, 0.61 mm broad. Measurements of the preabdominal sclerites: T3 15 × 38, T4 16 × 35, T5 19 × 33, T6 12 × 21, S2 18 × 29, S3 20 × 38, S4 20 × 36, S5 21 × 28, S6 14 × 20. Seventh and eighthscleritesverysmall, postabdomenmoderatelyextendable (abdominallengthc. 1.12 mm, additionalextendablepartc. 0.75 mm). T8 seeminglyin 3 parts (actuallynotso, areas betweendarkpartsonlyunpigmented). Sternite 8 minute, 0.05 mmlong. Femaleepiproct 0.08 × 0.04 mm, its setal pair 0.03 mm long. Hypoproct 0.06 mm broad with 2 pairs of submarginalsetae. Cercus 0.02 mmbroad, 0.06 mmlong, longestseta 0.08 mm, 2 mediumlongapicalandother 3 shortercercalsetaepresent. Spermathecaeovoid, c. 0.04 × 0.05 mm, ductsextremelyshort, only 0.012 mm .

Etymology. The specific epithet in the name of this new species refers to its type local- ity ( Morocco).

Remark. ItwasasurprisetofindanewspeciesfromthePalaearcticpart ofNorthAfrica. Ithasnopresuturaldcsetae (only 1 prescutellarpair) andits costalsetaearenormal; thiscombinationwouldbeenoughtoseparateitfrom theknownPalaearcticspecies. Inaddition, thearmatureofitsmalesternite 5 isunique: thestrongsetaeandpegsmakeitrecognizableinmostcaseseven withoutapreparationofthemaleabdomen).

Figures

Figs 78–83. Ceroptera moroccana sp. n., male, sternite 5 and genitalia: 78 = sternite 5, ventral view, 79 = anteriorlobeofleftsurstylus, broadest (sublateral) view, 80 = posteriorlobeof leftsurstylus, broadest (sublateral-subcaudal) view, 81 = phallusandphallapodeme, lateral view, 82 = postgonite, lateralview, 83 = contoursofsubepandrialsclerite, inner (anterior)