Psilopsyche granda, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 122-125

Psilopsyche granda , new species

Figs 272–282

This species resembles P. molinai Navás from Chile. Psilopsyche granda is distinguished from P. molinai in having large body size; in the genitalia segment X is more slender in lateral view; and the dorsal gap between the preanal appendages are wider and more U-shaped.

Male (in alcohol). Body large, light brown, ventral part of body and dorsal warts ochraceous; forewing membrane light brown, with darker spots and reticulation (in alcohol). Ocelli absent. Tentorium slender in dorsal view, without vestigial dorsal arm; anterior arms narrow at middle; posterior arms short, ending in pair of large posterior tentorial pits; tentorial bridge separating anterior and posterior tentorial arms slender, slightly arching anterad; anterior tentorial arms producing into well developed frontogenal septum in lateral view, forming internal fold extending dorsally into circumantennal sclerite; wide extension present below each anterior tentorial pits. Internal fold of frontogenal septa visible on facial surface as frontogenal sutures above anterior tentorial pits, and clypeogenal sutures below tentorial pits. Facial groove pattern dominated by surface grooves of frontogenal septa near tentorium; frontogenal vertical grooves running closely posterad. Clypeogenal vertical grooves located ventrally of each anterior tentorial pit; running obliquely laterad, near, but not reaching subgenal processes. Clypeolabral groove visible as line separating freely hanging, smooth labrum and clypeus; with single, dominating clypeal mesal compact setose wart. Subantennal groove running vertically in narrow stripe between ocellar grooves and frontogenal compact setose wart invisible. Subocular grooves indiscernible. Vertex 2 times wider than long. Epicranial groove complete; frontal branch present, vestigial, forming short Y-shape suture in dorsal view just anteriorly of vertexal interantennal compact setose warts. Coronal groove complete, clearly visible along entire length of vertex. Antennal sockets located on elevated humps with antennal grooves. Occipito-postgenal grooves visible on vertex between occipital and postgenal setal warts. Postoccipital groove encircling foramen magnum, or occipital foramen, producing pair of postoccipital smooth lobes. Labrum apparently freely hanging, membranous, movable, pyriform, without visible setae. Mandibles membranous, almost indiscernible; lacinia forming short, broad, mesad-curving setose lobe. Pair of large, ovoid, frontogenal compact setose warts located obliquely, dominating on faces; other warts visible on face is ovoid, horizontally located, clypeal mesal compact setose wart below anterior tentorial pits, between clypeogenal grooves. Anterior area of vertex with pair of enlarged, nearly triangular, well separated vertexal medioantennal compact setal warts. Vertexal lateroantennal compact setose warts absent. Occipital compact setose warts representing largest setal structure on vertex, located obliquely, dominating on entire surface of vertex. Vertically elongate postgenal compact warts curving along posterior section of ocular grooves. Postgenal surface glabrous. Maxillary palp formula (I-V-III-IV-II); fist segments bearing apicomesal nodule with erect apical setae. Scapes longer than head. Pedicels much shorter than first flagella segment. Pronotum with 2 pairs setal warts; mesal pair elevated, circular, well separated by deep depression; lateral pair weakly elevated, nearly triangular. Mesoscutum with pair of small, drop-shaped, diffuse setose warts present on middle of segment, at end of median notal suture. Mesoscutum with pair of small, rounded setose warts located on middle of segment. Each proepisternum with medium-sized, vertically elongate, ovoid, setose warts. Precoxale with large, nearly round wart. Large, compact, setal wart present anteriorly on cervical sclerites; forming sclerotized surface on membranous part of neck, anteriorly tangential with cervical sclerite. Lateral cervical sclerites forming narrow anterior arm articulating anteriorly to back of head with occipital condyle above posterior tentorial pits, fusing to posterior cervical sclerites; posterior cervical sclerites forming large, triangular plate broadening posterad, reaching prothoracic episternum by posterior apex; articulated with mesad-directed rods, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by thin ventral intercervical sclerites fused to posterior sclerites. Dark cervical sclerite complex visible on pale, membranous neck. Leg claws symmetrical; spur formula 2, 4, 4; foreleg anterior spur 2 times longer than posterior spur; midleg posterior spurs 2 times longer than anterior spurs; hind leg anterior spurs 3 times longer than posterior spurs; all spurs with fine, serrated mesal edge; surfaces with microsculpture with few vestitural setae; spurs on all legs ochraceous. Legs covered by thin, short, light brown, vestitural setae, more densely on ventral surfaces, interspersed by erect and spine-like setae only on tarsal segments of mid and hind legs. Forewing narrow, length 22.0 mm; membrane light brown with spotted, reticulated darker pattern; termen slightly concave posteriorly; posterior margin concave; basal lobe covered by microtrichia; Sc running free before C; R1 running to R2 before C; postanal vein running near posterior wing margin.

Male genitalia. Abdominal Segment IX fused annularly; short in lateral view; tergum almost as long as venter; anterior margin slightly convex, almost straight; posterior margin concave, without apical lobe; antecosta and antecostal suture on anterior margin narrow, forming dark marginal rim running evenly along margin; entire surface of segment IX glabrous; spine row absent on posterior margins of segment IX; setose areas absent from apicopleural and apicoventral regions. Intersegmental depression between segment IX and segment X deep, stepwise. Segment X sunk deep to middle of segment IX, visible in lateral view. Segment X slender in lateral view, with 2 mesal, triangular apical lobes in dorsal view. Apicoventral setose lobes forming blunt ventral corner. Apicodorsal setose lobes long, tapering, obliquely directed dorsad; setose surface expanding over almost entire dorsal and ventral parts of segment. Apex of segment X deeply cleft; dorsal interlobular gap long, parallel-sided. Large preanal appendages dominating over phallic apparatus; horizontally posterad oriented lobes fused to dorsum of segment IX. Fused seam, or borderline between segment IX and preanal appendages, hardly visible; in lateral view, dorsum slightly convex, apex triangular. In dorsal view mesal margins of appendages running parallel in dorsal view. Coxopodites producing into elongate apicodorsal corners. Harpagones clavate in ventral view; with dark, short, conical spines on apical surfaces. Phallic apparatus forming sclerotized phallotheca with ventral lobe and of membranous apical endotheca and retracted phallicata. Sclerotized phallotremal sclerites complex, retracted basad, consisting of basoventral, single tube-like elongate vesicle, and apicodorsal paired wing-shaped sclerites. Thin ejaculatory duct forming entering vesicle.

Holotype male: CHILE: Region del Araucania ( IX): PN Nahuelbuta, camp site, Estero Cabreria, crosspoint between streams draining Mts Pichimanquemáhuida and Pichinahuel , 37°49.647’S, 73°00.691’W, 1100 m (loc#07), light trap 3.i.2006 [K.A. Johanson] ─ ( NRM) . Paratypes: same data as holotype — 6 males ( NRM), 2 males ( OPC). Distribution: Chile. Etymology: granda , from Latin, large, referring to the large size of this species .

— genitalia, dorsal; 280 — genitalia, ventral; 281 — phallus, lateral; 282 — phallus, ventral.

Figures

FIGURES 272–276. Psilopsyche granda, new species, holotype. 272 — head, frontal; 273 — head dorsal; 274 — hear, lateral; 275 — tentorium, dorsal; 276 — tentorium, lateral.

FIGURES 277–282. Psilopsyche granda, new species, holotype. 277 — right wings; 278 — genitalia, lateral; 279