Magnanillus regalis, Giachino & Eberhard & Perina, 2021

Giachino, Pier Mauro, Eberhard, Stefan & Perina, Giulia, 2021, A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia, ZooKeys 1044, pp. 269-337 : 269

Magnanillus regalis sp. nov. Figs 24-26

Type locality.

WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings deposit, 22°07'45.5"S, 117°52'24.3"E.

Type series.

HT ♂, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings Mine, 22°07'45.5"S, 117°52'24.3"E (WGS84), G. Pearson and D. Main, 12 Jan 2010. Trog. net scrape (SM3175), Western Australian Museum Entomology Reg. no. 82653 (WAM). PTT: 1 ♂ 2 ♀♀ (remains), WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings Mine, 22°07'45.5"S, 117°52'24.3"E (WGS84), G. Pearson and D. Main, 12 Jan 2010. Trog. net scrape (SM3175), Western Australian Museum Entomology Reg. no. 82653 (WAM, CGi).

Differential diagnosis.

Magnanillus regalis sp. nov. is easily distinguishable from M. firetailianus sp. nov. and M. quartermaini (Baehr & Main, 2016) by its pronotum with basal border ca. as wide as the anterior border. It can be distinguished from M. sabae sp. nov. by its longer metatrochanters, reaching the femoral tooth. It differs from M. serenitatis sp. nov. by its shorter metatrochanters, not overreaching the femoral tooth. It can be distinguished from M. salomonis sp. nov. by its less transverse pronotum and the straight apex of its metatrochaters.

Description of the HT ♂.

TL mm 2.25. Body elongate, depigmented, yellow-testaceous; integument shiny with evident microsculpture and short pubescence.

Head relatively large, narrower than pronotum; with two couples of excess setae on the vertex as in Fig. 24 . Labium without tooth, mentum articulated. Antennae robust, submoniliform, very short, not reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum sub-squared (max. width / max. length ratio = 1.14 maximum width at the base of the anterior fourth, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in anterior part, subrectilinear at the basal half, not sinuate but with an evident tooth before basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly placed internally on the disk and before the posterior angles.

Legs long and slender, with metatrochanters long, acuminate, subrectilinear and metafemora dentate; metatrochanters (Fig. 25 ) reaching the femoral tooth. Two dilated protarsomeres, without adhesive phanerae in males.

Elytra subrectangular, elongate (max. length / max. width ratio = 1.83), not truncated and only slightly emarginated before apex. Disc convex, with longitudinal grooves; integument shiny, with evident microsculpture and very short, longitudinally aligned, upright pubescence. Humeri well marked, obtuse; post-humeral margin denticulate, with distinct crenulations up to the base of the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8th pore of the umbilicate series.

Chaetotaxy: scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4th pore farther and placed at the end of the basal third of the elytron; 5th pore placed at the base the apical third of the elytron; 7th pore very forward, placed near the 6th pore; 6th and 7th pores closer than 5th and 6th; 8th displaced onto the disc; 7th and 8th spaced out ca. the 5th and 6th. Three discal setae, first placed before the 4th pore of the umbilicate series, second one placed just before the 5th pore, third one placed after the 7th pore.

Aedeagus (Fig. 26 ) large, median lobe long, slender, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, short. Endophallus without an evident lamella copulatrix, but with small, Y-shaped, apical, slightly sclerified stripe. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.

Etymology.

The name comes from the Latin word Magnanillus regalis = royal, and it reminds the type locality “King” deposit in the Solomon Mining Area.

Distribution.

Magnanillus regalis sp. nov. is known only from the type locality (Kings deposit) in the Solomon Mining Area, 50 km N of Tom Price, Pilbara, WA.

Figures

Figures 24 - 26. Magnanillus regalis sp. nov., HT ♂ 24 habitus 25 right metafemur and metatrochanter in ventral view 26 aedeagus in lateral view. Scale bars: 0.1 mm.