Ummidia varablanca, Godwin & Bond, 2021

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia varablanca sp. nov. Figs 59 , 60 , Map 5

Type material.

HOLOTYPE: 1 ♂ (UMM669) from 9 kilometers east of Vara Blanca transect, Heredia, Costa Rica, 10.0024 -84.1216 1, 1152 m a.s.l., 8.iii.2005, BME. PARATYPE: 1 ♀ (UMM746) from Alto Tapezco, San José, Costa Rica, 9.8982 -84.1553 1, 2.viii.2001, coll. MA Zumbado, BME.

Etymology.

The specific epithet is a noun taken in apposition and is in reference to the type locality, Vara Blanca.

Diagnosis.

Ummidia varablanca are relatively small spiders (CL 3-4.8) with distinctly light tarsi. Males can be distinguished from U. hondurena , U. yojoa , U. rugosa , and U. erema by having a relatively short embolus with a single even curve and from all except U. zebrina and U. yojoa by lacking prolateral spines and having very few retrolateral spines (2 vs 10-44) on tibia I. Males can be further differentiated from U. rugosa and U. riverai by the lack of a pale dorsal heart patch on the abdomen. Females can be differentiated from all geographically proximate species by having spermathecae which curve medially then bend strongly anteriorly.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. General coloration. Carapace, chelicerae, and legs reddish black 2.5YR 2.5/1, tarsi brownish yellow 10YR 6/8. Abdomen black 5YR 2.5/1. Cephalothorax. Carapace 3.35 long, 3.46 wide. Pars cephalica 2.44 long. Foveal groove procurved, 0.3 long, 0.6 wide. All eyes on defined tubercle. AER procurved. PER straight. Eye group 0.53 long, 1.03 wide, AME 0.27, PME 0.18, ALE 0.3, PLE 0.19. Sternum sparsely setose, STRl 1.94, STRw 2.05. Chelicerae with anterior tooth row comprising three teeth, posterior margin with five teeth. Palpal endites with mix of 21 small and large, hastate and non-hastate cuspules spread over proximal 4/5 of endite face, lacking distal endite cuspules, ENDw 0.71, ENDl 1.21. Labium with five cuspules, LBw 0.8, LBl 0.51. Rastellum with six fine spines along distal margin. Abdomen setose. Legs. F1 3.37; F1w 0.74; P1 1.66; Ti1 2.2; Mt1 1.65; Tr1 0.9; F3 2.54; F3w 0.88; P3 1.22; Ti3 1.63; Sd3 0.92; Mt3 1.4; Tr3 1.01; F4 3.16; F4w 0.83; P4 1.48; Ti4 1.97; Mt4 1.86; Tr4 0.98. Retrolateral face of tarsus IV without brush or comb. Leg I spination pattern: TSp 0, TSpv 0, TSrd 0, TSr 0, TSrv 2, MtSp 0, MtSr 0, TrSp 0, TrSr 0. Pedipalps. PTl 1.8, PTw 0.64, Bl 1.23. Embolus evenly curved.

Variation, males

(n = 7). CL 2.99-4.04, 3.45 ± 0.12; CW 3.04-3.96, 3.5 ± 0.11; Cap 2.08-2.82, 2.38 ± 0.09; ENDl 0.46-0.63, 0.54 ± 0.02; ENDw 0.83-1.19, 0.99 ± 0.05; STRl 1.72-2.21, 1.93 ± 0.06; STRw 1.64-2.17, 1.94 ± 0.06; LBl 0.44-0.58, 0.51 ± 0.02; LBw 0.67-0.85, 0.78 ± 0.02; F1 2.82-3.87, 3.26 ± 0.12; F1w 0.72-0.9, 0.81 ± 0.02; P1 1.38-1.8, 1.56 ± 0.05; Ti1 1.75-2.47, 2.03 ± 0.1; Mt1 1.2-1.86, 1.47 ± 0.09; Tr1 0.73-1.74, 0.94 ± 0.14; F3 2.07-2.81, 2.42 ± 0.09; F3w 0.78-1.04, 0.88 ± 0.03; P3 1.03-1.37, 1.18 ± 0.04; Ti3 1.24-1.77, 1.5 ± 0.08; Mt3 1.12-1.5, 1.33 ± 0.06; Tr3 0.78-1.09, 0.96 ± 0.04; F4 2.71-3.66, 3.07 ± 0.12; F4w 0.74-0.93, 0.84 ± 0.02; P4 1.18-1.55, 1.37 ± 0.05; Ti4 1.64-2.27, 1.87 ± 0.08; Mt4 1.67-2.24, 1.87 ± 0.08; Tr4 0.73-1.01, 0.93 ± 0.04; TSp 0-8, 1.71 ± 1.13; TSpv 0-6, 3 ± 0.79; TSr 0-2, 0.57 ± 0.3; TSrv 2-19, 10 ± 1.9; PTl 1.28-2.09, 1.66 ± 0.1; PTw 0.59-0.75, 0.66 ± 0.02; BL 1.23-1.53, 1.34 ± 0.04.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. General coloration. Carapace, chelicerae, and legs very dark brown 7.5YR 2.5/3, TrIV strong brown 7.5YR 5/8. Abdomen black 7.5YR 2.5/1, spinnerets dark yellowish brown 10YR 3/6. Cephalothorax. Carapace 4.03 long, 3.81 wide. Pars cephalica 3.07 long. Foveal groove procurved, 0.45 long, 0.96 wide. All eyes on moderate tubercle. AER procurved. PER straight. Eye group 0.6 long, 1.04 wide, AME 0.25, PME 0.2, ALE 0.36, PLE 0.27. Sternum sparsely setose, STRl 2.25, STRw 2.45. Chelicerae with anterior row comprising five teeth, posterior margin with six teeth. Palpal endites with 37 large cuspules across proximal 3/4 of endite face and ten smaller cuspules distally, ENDw 0.91, ENDl 1.56. Labium with seven large cuspules, LBw 0.98, LBl 0.64. Rastellum reduced with long rounded spines along margin. Abdomen. Evenly setose with pale speckles clustered at apodemes. Legs. F1 2.66; F1w 0.99; P1 1.58; Ti1 1.52; Mt1 1.08; Tr1 0.74; F3 2.37; F3w 1.15; P3 1.24; Ti3 1.27, Sd3 0.89; Mt3 0.93; Tr3 0.84; F4 2.86; F4w 1.15; P4 1.38; Ti4 1.51; Mt4 1.41; Tr4 0.9. Retrolateral face tarsus IV with comb of alternating long and short spinules. Pedipalps. PF 2.52, PP 1.34, PTi 1.38, PTr 1.38. Spermathecae curved medially and then bent anteriorly, bulbs facing anteriorly.

Variation, females

(n = 3). CL 3.96-4.83, 4.27 ± 0.28; CW 3.81-4.32, 4.07 ± 0.15; Cap 2.82-3.42, 3.1 ± 0.17; ENDl 0.6-0.66, 0.64 ± 0.02; ENDw 1.04-1.15, 1.08 ± 0.04; STRl 2.25-2.82, 2.51 ± 0.17; STRw 2.45-2.6, 2.52 ± 0.04; LBl 0.64-0.73, 0.67 ± 0.03; LBw 0.98-1.1, 1.03 ± 0.04; F1 2.66-3.12, 2.87 ± 0.13; F1w 0.99-1.12, 1.04 ± 0.04; P1 1.58-1.88, 1.73 ± 0.09; Ti1 1.52-1.77, 1.61 ± 0.08; Mt1 1.08-1.22, 1.13 ± 0.04; Tr1 0.74-0.86, 0.78 ± 0.04; F3 2.37-2.68, 2.52 ± 0.09; F3w 1.15-1.32, 1.23 ± 0.05; P3 1.24-1.42, 1.36 ± 0.06; Ti3 1.27-1.38, 1.34 ± 0.03; Mt3 0.93-1.1, 1 ± 0.05; Tr3 0.84-1.1, 0.95 ± 0.08; F4 2.86-3.45, 3.07 ± 0.19; F4w 1.15-1.37, 1.22 ± 0.07; P4 1.38-1.79, 1.58 ± 0.12; Ti4 1.51-1.88, 1.67 ± 0.11; Mt4 1.41-1.65, 1.53 ± 0.07; Tr4 0.9-1.09, 0.98 ± 0.06; PF 2.52-2.82, 2.65 ± 0.09; PP 1.34-1.63, 1.46 ± 0.09; PTi 1.38-1.6, 1.5 ± 0.07; PTr 1.38-1.67, 1.48 ± 0.1.

Material examined.

Costa Rica: Cartago: El Copal, Pejibaye , 9.7838 -83.7489 1, 992 m a.s.l. (UMM0751, vi.2006, 1♀, C Viquez, BME); Esperanzo del Gaurco, Tapanti National Park, Biological Station, 9.7585 -83.7843 1, 1261 m a.s.l. (UMM0668, 3.v.2002, 1♂, BME) ; Guanacaste: Rio San Lorenzo, Tierras Morenas , 10.569 -85.0206 1, 1500 m a.s.l. (UMM0681, 1995, 1♂, G Rodriguez, BME) ; Heredia: Heredia, 6 km ENE of Vara Blanca transect, 10.0024 -84.1216 1, 1152 m a.s.l. (UMM0670, 9.iv.2002, 1♂, BME); Heredia, 9 km E of Vara Blanca transect, 10.0024 -84.1216 1, 1152 m a.s.l. (UMM0669, Summer 2005, 1♂, BME); Vara Blanca , 10.1678 -84.1475 1, 1876 m a.s.l. (UMM0750, 16.iii.2002, 1♀, BME) ; Puntarenas: Agujas, 8.5137 -83.5453 1, 85 m a.s.l. (UMM0675, 1♂, BME); Monteverde, El Valle de Gutierrez , 10.299 -84.803 1, 1600 m a.s.l. (UMM0673, 5-7.x.2009, 1♂, BME); Monteverde, San Luis, Buen Amigo , 10.2721 -84.824 1, 1300 m a.s.l. (UMM0674, ii.1995, 1♂, Z Fuentes, BME) ; San José: Alto Tapezco , 9.8982 -84.1553 1, 1615 m a.s.l. (UMM0746, 1♀, BME) .

Figures

Map 5. Locality records for southern Mexico and Central American species.

Figure 59. Ummidia varablanca sp. nov. from Heredia, Costa Rica. A male habitus illustration UMM 669 B female habitus illustration UMM 746. Scale bars: 4.0 mm.

Figure 60. Ummidia varablanca sp. nov. from Heredia, Costa Rica A-D male holotype (UMM 669) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 746). Scale bars: 1.0 mm.