Phlegmariurus foliaceus ­­­ (Maxon) B. Øllgaard (2012b: 15)

Øllgaard, Benjamin & Testo, Weston, 2021, The Lycopodiaceae of Panamá, Phytotaxa 526 (1), pp. 1-66 : 23-26

. ­­­ Phlegmariurus foliaceus ­­­ (Maxon) B. Øllgaard (2012b: 15) . Fig. 9

Lycopodium foliaceum Maxon (1912: 1) .— Urostachys foliaceus (Maxon) Nessel (1939: 68) .— Huperzia foliacea (Maxon) Holub (1985: 72) . TYPE:— PANAMÁ, Chiriquí: Humid forest along the upper Caldera River, near »Camp 1« Holcomb’s trail, above El Boquete, 1450–1650 m, 22–24 Mar 1911, Maxon, W. R. 5628 (AAU, GH, US).

Erect from an ascending base, up to 50 cm tall, sparsely branched. Shoots homophyllous, almost equally thick throughout, 15–30 mm in diam. incl. leaves, sometimes tapering upward. Stems excl. leaves 2–4 mm thick near the base, somewhat ridged by decurrent leaf bases. Leaves uniform throughout, borne in alternating whorls of 4–5, these 2–4 mm apart, wide-spreading to somewhat reflexed, usually straight, not twisted at the base, oblong-elliptic to oblanceolate from a narrow long-cuneate base, apiculate, 6–17 × 1.5–3.5 mm, almost flat, with prominent vein above, or folded slightly down along the vein, papery to subcoriaceous, with slightly revolute, apically denticulate margins. Leaf bases often red-dotted; sporangia (1.5–) 2 (–2.5) mm wide.

­­­ Distribution:— Costa Rica, Panamá. Narrowly endemic.

­­­ Habitats:—Epiphytic in wet forest. 1050–1900 m elev.

­­­ Specimens­­­studied:— Bocas­­­del­­­Toro:Forest near headwaters of Río Culebra, ca. 5 km ENE of Cerro Pate Macho, E of Finca Serrano, 1160 m, Hammel 6145 (MO). Chiriquí: Upper Caldera river, near Camp I, Holcomb’s Trail, above El Boquete, 1450–1650 m, Maxon 5623a ( US), 5628 (AAU, C, US). Along Road Gualaca–Fortuna Dam site, at 10.1 mi NW of Los Planes de Hornito, 1260 m, Antonio 4195 (MO). Road Fortuna Lake–Chiriquí Grande, 4.5–5 km N of dam over lake, 1100–1135 m, Croat 59945 (MO). La Fortuna Hydroelectric project, 1200 m, Hammel 2195 (MO). Fortuna Dam region, Along Quebrada Arena, 1050 m, McPherson 8733 (MO). Bajo Chorro, near Río Caldera, 1900 m, Skog et al. 4064 (MO, US). Near Boquete, trail from Bajo Chorro along the Río Caldera, 2.5 km from trailhead, 1700 m, W. Testo 1000 (PMA, VT).