Coenosia luxia, Sorokina, 2023

Sorokina, Vera S., 2023, A key to the Russian species of the genus Coenosia Meigen (Diptera, Muscidae), with the description of one new species and new synonymies, Zootaxa 5389 (1), pp. 79-107 : 100-104

Key to females

The females of C. flavissima , C. kosterini , C. rhaensis , C. sibirica and C. striolata are not known, but the last two species are included in the key according to the characters of the males.

1. Costa extending only to, or very slightly beyond, apex of R 4+5 .................................................. 2

- Costa extending to apex of M 1........................................................................... 3

2. Lower calypter not longer than upper one. Postpedicel with a spine-like dorsal tip.................. connectens (Hennig)

- Lower calypter much longer than upper one. Postpedicel without a spine-like dorsal tip, with a more or less rounded tip........................................................................................ agromyzina (Fallén)

3. Hind tibia with a median anterior seta and an anterodorsal seta at almost the same level and close together ( Fig. 9A )....... 4

- Hind tibia without an anterior seta at middle close to the anterodorsal seta, but an anteroventral seta present............. 9

4. Scutum, scutellum and abdomen shining black.................................................... atra (Meigen)

- Scutum, scutellum and abdomen not shining black........................................................... 5

5. Parafacial at least as broad as postpedicel; longest aristal hairs at least as long as width of postpedicel; outer vertical setae longer and stronger than postocular setulae......................................................... tigrina (Fabricius)

- Parafacial narrower than width of postpedicel; longest aristal hairs much shorter than width of postpedicel; outer vertical setae at most as long as postocular setulae...................................................................... 7

7. All femora yellow, more or less darkened in apical third; abdomen with a midline and paired brown spots.... strigipes Stein

- All femora dark, at most the tips narrowly yellow; abdomen with or without paired spots............................ 8

8. Longest aristal hairs barely longer than basal width of arista; abdomen at most with indistinct pattern.—European Russia........................................................................................... attenuata Stein

- Longest aristal hairs at least half as long as width of flagellomere; abdomen with paired round spots....... humilis Meigen

9. Hind tibia with strong and long preapical dorsal and posterodorsal setae close to each other ( Fig. 9B ).................. 10

- Hind tibia without strong and long, closely approximated preapical dorsal and posterodorsal setae.................... 11

10. Mid and hind femora and all tibiae predominantly yellow...................................... intermedia (Fallén)

- Mid and hind femora and all tibiae predominantly dark but fore and mid trochanters yellow, hind trochanter sometimes darker.—European Russia................................................................... means Meigen

11. Head almost square in lateral view ( Fig. 10F ); frons reddish at antenna, postpedicel black, very long, 4 times as long as wide; oral vibrissae usually yellow or golden but sometimes black...................................... flaviseta Huckett

- With a different combination of characters................................................................ 12

12. Lower calypter shorter than the upper one................................................................. 13

- Lower calypter similar to or longer than upper one.......................................................... 16

13. Basal scutellar setae reduced, apical setae weak and short ( Fig. 7B )................................... ciliata Hennig

- Both pairs of scutellars strong and long................................................................... 14

14. Larger size: 6–7.5 mm ( Fig. 10B ). Mid and find femora completely yellow; mid femur with a strong anterior preapical seta............................................................................... baicalensis Schnabl (part)

- Smaller size: 3.5–5.5 mm. Mid and find femora completely or partially dark; mid femur without a strong anterior preapical seta............................................................................................... 15

15. At least tibiae yellow ( Fig. 10C ); palpus at least yellow on basal half; lower calypter extremely small, shorter than half length of the upper one, strip-like; legs without hairs on ventral surface; preapical anterodorsal and posterodorsal setae on hind tibia indistinct or absent....................................................................... tendipes Huckett

- All tibiae black, only knees yellowish; palpus black; lower calypter as broad as half length of the upper one; legs with hairs on ventral surface; preapical anterodorsal seta on hind tibia long and strong............................ penicillata Hennig

16. Lower calypter subequal in length or scarcely longer than upper one............................................ 17

- Lower calypter much longer than upper one............................................................... 34

17. All femora black, at most with apices narrowly yellow....................................................... 18

- At least mid and hind femora yellow on basal 1/3........................................................... 25

18. Antenna including both basal segments yellow or at least basal segments and base of postpedicel yellowish................................................................................................. dealbata (Zetterstedt)

- Antenna entirely black................................................................................ 19

19. All tibiae predominantly black, at most yellow in basal third.................................................. 20

- All tibiae yellow or brownish-yellow..................................................................... 23

20. Mid femur with a strong distinct anterior preapical seta; scutum with 2 distinct brown narrow longitudinal vittae along dorsocentral setae; scutellum grey, sometimes like scutum with yellowish dust ( Fig. 7E )........... subgracilis Xue & Cui

- Mid femur without anterior preapical seta or with indistinct short seta; scutum with brown dust between dorsocentral setae, reaching the apex of scutellum or completely brown dusted................................................... 21

21. Gena and parafacial broad, height of gena 2 times width of postpedicel, width of parafacial about as wide as postpedicel; mid femur usually with apical posteroventral seta (Figures in Sorokina 2022).................. wrangelensis Sorokina (part)

- Height of gena 1.5–1.7 times width of postpedicel, width of parafacial at middle as wide as ½ or 2/3 of width of postpedicel; mid femur without apical posteroventral seta.............................................................. 22

22. Scutum completely brown dusted; hind femur with a row of short anteroventral setae (Figures in Sorokina 2022)................................................................................................. galina Sorokina

- Scutum with thick brown dust between dc which reaches the apex of scutellum; hind femur with two long anteroventral setae, which are much longer then diameter of femur in distal part and with two anteroventral setae in proximal part of femur (Figures in Sorokina 2022)..................................................................... koni Sorokina (part)

23. Mid femur without a strong anterior preapical seta; preapical anterodorsal and posterodorsal setae on hind tibia distinct, each half as long as preapical dorsal seta. Smaller species (3.5–4.7 mm) ( Fig. 10H ).................. conflicta Huckett (part)

- Mid femur with a strong anterior preapical seta; preapical anterodorsal on hind tibia much longer and stronger than posterodorsal setae. Larger species (5.0– 6.5 mm)...................................................................... 24

24. All tibiae yellow; abdomen with rounded spots; mid femur with a full row of anteroventral setae ( Fig. 7G )....................................................................................................... oralis Schnabl

- All tibiae brownish-yellow, darkened in apical 2/3–4/5; abdomen with more or less triangular spots; mid femur with 5–6 anteroventrals in basal half ( Figs 7I , 8F ).............................................. ukokensis Sorokina (part)

25. Mid femur in apical third without a strong anterior preapical seta.............................................. 26

- Mid femur with a strong anterior preapical seta............................................................. 28

26. Body entirely yellow; scutum densely grey dusted, pleura yellow with grey dusted spot on anepimeron; apical tarsomeres of all legs often blackish.—Far East (Kunashir Island)................................................. polina Vikhrev

- Al least thorax and abdomen grey, not yellow.............................................................. 27

27. Scutum with two distinct brown longitudinal stripes; antenna and palpus wholly dark; fore femur dark, mid and hind femora yellow or with dark dorsal streak.................................................. bilineella (Zetterstedt) (part)

- Scutum brownish dusted, without dark longitudinal stripes; at least base of postpedicel yellow; palpus partly yellow or brownish; all femora yellow or fulvous............................................................... demoralis Huckett

28. Fore femur mostly black, mid and hind femora black or fuscous in dorsal part at apex; antenna completely black........ 29

- All femora completely yellow, at most fore femur with dark dorsal steak; antenna at least yellow at base on ventral surface (except black in C. baicalensis )......................................................................... 31

29. Fore femur darkened at most on dorsal surface, ventral surface yellow; mid and hind femora largely yellow on distal half with a dark mark at most confined to dorsal half of femur; mid femur with 1 preapical posterior seta.... cingulipes (Zetterstedt)

- Fore femur completely black; mid and hind femora with a black apical ring; mid femur with 2 preapical posterior setae... 30

30. Hind tibia with a well-developed posterodorsal seta, as long as 2/3 of anterodorsal seta; gena higher than width of postpedicel ( Fig. 9F )............................................................................. alaskensis Huckett

- Hind tibia without posterodorsal seta, or with some a weak and short seta which shorter than anterodorsal seta; gena not higher than width of postpedicel.......................................................... octopunctata (Zetterstedt)

31. Antenna and palpus wholly dark; mid and hind femora with complete but sparse rows of long posteroventral setae; height of gena 1.5 times width of postpedicel ( Fig. 10B ). Larger size: 6–7.5 mm ...................... baicalensis Schnabl (part)

- Antenna and palpus partly or completely yellow; mid and hind femora with long posteroventrals only in basal half; height of gena at most as wide as width of postpedicel. Smaller size: 3–5 mm ............................................ 32

32. Postpronotal lobe, abdomen and at least katepisterna yellow..................................... ghilarovi Lobanov

- Thorax completely grey, abdomen yellow only at base....................................................... 33

33. Tip of antennal pedicel yellow; palpus at least broadly yellow at tip, usually almost or completely yellow; dark dorsocentral vittae weakly visible; fore femur wholly yellow............................................... mollicula (Fallén)

- Antennal pedicel and postpedicel usually narrowly reddish basally; palpus wholly dark or at least dark at tip; dark dorsocentral vittae conspicuous and strongly marked; fore femur yellow or with a dark dorsal steak............... alpicola (Pokorny)

34. Postpedicel with a spine-like dorsal tip ( Fig. 9D )........................................................... 35

- Postpedicel without a spine-like dorsal tip, rounded at tip..................................................... 38

35. Mid and hind femora yellow but darkened apically............................................. ambulans Meigen

- Mid and hind femora predominantly black................................................................ 36

36. Height of gena 1.5–1.8 times width of postpedicel ( Fig. 1C ); legs completely black, grey dusted like pleura; two proepisternal setae..................................................................................... luxia sp. nov.

- Height of gena less than 1.5 times width of postpedicel, usually as high as wide of postpedicel; at least knees and base of fore and mid tibiae yellow; usually one proepisternal setae....................................................... 37

37. Frons uniformly silvery; abdominal spots brown, well defined................................ flavimana (Zetterstedt)

- Frontal vitta darker than fronto-orbital plates; abdominal spots indistinct........................................................................................... pulicaria (Zetterstedt) & acuminata Strobl (European Russia)

38. Only one proepisternal seta (sometimes with an additional weak seta on one side)................................. 39

- Two proepisternal setae............................................................................... 45

39. All femora black, at most only narrowly yellow apically..................................................... 40

- Femora completely yellow or yellow in apical third......................................................... 41

40. All tibiae yellow ( Fig. 10H ); scutum grey dusted with 2 or 3 brownish vittae, scutellum grey dusted; frontal vitta brownishgrey dusted, fronto-orbital plate silvery-grey dusted; preapical anterodorsal and posterodorsal setae on hind tibia half as long as length of preapical dorsal seta........................................................ conflicta Huckett (part)

- All tibiae black; scutum and scutellum completely densely brown dusted, sometimes with 3 indistinct longitudinal vittae; frontal vitta and a fronto-orbital plate velvety black in frontal view; preapical anterodorsal seta on hind tibia as long as 2/3 length of preapical dorsal seta, preapical posterodorsal absent (Figures in Sorokina 2022)............... adriani Sorokina

41. Postpedicel shorter, ca. 2.2–2.3 times as long as wide. Mid tibia with apical posteroventral seta short, not longer than diameter of tibia where situated.............................................................. incisurata van der Wulp

- Postpedicel longer, ca. 3.0 times as long as wide. Mid tibia with apical posteroventral seta longer or not than diameter of tibia where situated....................................................................................... 42

42. Lower calypter short, little longer than upper one; frontal vitta entirely dark; mid tibia with apical posteroventral seta longer than diameter of tibia where situated............................................... bilineella (Zetterstedt) (part)

- Lower calypter much longer than upper one; anterior margin of frontal vitta reddish-yellow or dark; mid tibia with apical posteroventral seta shorter than or as long as diameter of tibia where situated..................................... 43

43. All coxae yellow; abdomen without or with very indistinct dark paired spots; presutural acrostichal setae weak................................................................................................. rufipalpis Meigen

- Mid and hind coxae partly grey; abdomen with distinct dark paired spots; presutural acrostichals well developed........ 44

44. Anterior margin of frontal vitta reddish-yellow; mid femur with only 1–2 weak posteroventrals in basal 2/3 and mostly with one apical posterodorsal seta; mid tibia with apical posteroventral seta much shorter than diameter of tibia where situated.................................................................................. testacea (Robineau-Desvoidy)

- Anterior margin of frontal vitta usually dark; mid femur with 3–4 posteroventrals in basal 2/3 and usually with 2 apical posterodorsals; mid tibia with apical posteroventral seta ca. as long as diameter of tibia where situated....................................................................................................... nigridigita Rondani

45. Hind tibia with one strong or 3–4 short submedian posterodorsal seta........................................... 46

- Hind tibia without a submedian posterodorsal setae......................................................... 55

46. Middle and hind femora black at least in distal half or entirely black; 2 pairs of strong presutural dorsocentral setae present (except 1 pair in C. ozerovi )............................................................................ 47

- Femora completely or predominantly yellow; only one pair of presutural dorsocentrals............................. 51

47. Middle and hind femora yellow in basal part............................................................... 48

- Middle and hind femora entirely black................................................................... 49

48. Middle and hind femora yellow in basal ½ part ( Fig. 11E ); hind tibia usually with 1 strong posterodorsal seta..................................................................................................... xuei Cui et Li

- Middle and hind femora yellow in basal ¼ or 1/3 ( Fig. 11D ); hind tibia with 3–4 short hair-like posterodorsal setae....................................................................................... apukaensis Hennig (part)

49. One pair of strong presutural dorsocentral setae present; all tibiae black, knees yellow; abdomen without dark spots ( Fig. 8B )....................................................................................... ozerovi Vikhrev

- Two pair of strong presutural dorsocentral setae present; tibiae yellow at least in basal third; abdomen with dark spots on tergites 3–5......................................................................................... 50

50. Scutum with 3 brown longitudinal vittae; lower calypter much longer than upper one; postpedicel 3.5 times as long as wide, nearly reaching vibrissal angle ( Fig. 8C ).................................................... nigrotincta Hennig

- Scutum with 2 brown longitudinal vittae; lower calypter slightly longer than upper one; postpedicel 2.5 times as long as wide, its length about 2/3 height of face ( Figs 7I , 8F )......................................... ukokensis Sorokina (part)

51. Postpedicel yellow basally............................................................... flavicornis (Fallén)

- Postpedicel completely black........................................................................... 52

52. Mid femur in distal third without long posteroventral setae.................................. trilineella (Zetterstedt)

- Mid femur in distal third with strong posteroventrals........................................................ 53

53. Postpedicel long, about 5.0 times as long as wide, reaching the mouth margin; parafacial very narrow, much narrower than width of postpedicel; fore femur with brown dorsal streak; mid and hind femora darkened apically; arista short pubescent....................................................................................... emiliae Lukasheva

- With a different combination of characters................................................................ 54

54. Posterodorsal seta of hind tibia as long and strong as the anterodorsal seta ( Fig. 12B ).................. lacustris Schnabl

- Posterodorsal seta of hind tibia shorter than or even half as long as the anterodorsal ( Fig. 12D ).......... luteipes Ringdahl

55. Apical scutellar setae distinctly shorter and weaker than basal scutellar setae..................................... 56

- Apical and basal scutellars subequal in length.............................................................. 57

56. Mid and hind femora black; mouth margin projecting slightly beyond level of profrons.................. pedella (Fallén)

- Mid and hind femora yellow; mouth margin not projecting beyond level of profrons............... griseiventris Ringdahl

57. Two pairs of presutural dorsocentral setae, anterior pair about 1/3–2/3 as long as posterior pair ( C. apukaensis Hennig sometimes without distinct anterior pair)........................................................................... 58

- Only one pair of presutural dorsocentrals................................................................. 62

58. At least base of postpedicel yellowish; base of abdomen more or less yellow.................... ruficornis (Macquart)

- Antenna entirely black; abdomen entirely grey............................................................. 59

59. At least basal third or quarter of mid and hind femora yellow; hind femur with a full row of strong posteroventrals....................................................................... comita (Huckett) & apukaensis Hennig (part)

- All femora black, only narrowly yellow apically; hind femur with strong posteroventral setae only on basal half......... 60

60. All tibiae yellow. Larger size: 6.0–8.0 mm.......................................... Macrorchis meditata (Fallén)

- At least hind tibia black. Smaller size: 3.5–4.3 mm .......................................................... 61

61. Fore and mid tibiae yellow, hind tibia black; hind tarsi thickened, all segments about twice as wide as those of mid tarsus; height of gena ca. 1.3 times width of postpedicel ( Fig. 11B )...................................... atritibia Ringdahl (part)

- All tibiae black, except yellow basal third or fourth; hind tarsi not thickened; height of gena ca. 2.3–3.0 times width of postpedicel (Figures in Sorokina 2022)......................................................... gorodkovi Sorokina (part)

62. Mid and hind femora predominantly yellow, darkened dorsally at apex.......................................... 63

- Mid and hind femora predominantly black, basal half never entirely yellow...................................... 69

63. Anteroventral seta on hind tibia in strictly ventral position......................................... pudorosa Collin

- Anteroventral seta on hind tibia in a normal, truly anteroventral position......................................... 64

64. Smaller size, the largest 3.5 mm. Mid and hind femora completely yellow or with dark dorsal spot at apex; hind femora with posteroventral setae only in basal half.................................................................... 65

- Larger size, more than 3.5 mm. Mid and hind femora yellow with black rings in apical fourth; hind femora with posteroventral setae in basal 2/3..................................................................................... 66

65. Mid and hind femora completely yellow, without dark dorsal spot at apex........................ mandschurica Hennig

- Mid and hind femora yellow with dark dorsal spot at apex......................................... dubiosa Hennig

66. Postpedicel 3.0 times as long as wide; scutum with 3 dark longitudinal vittae; presutural acrostichals in one irregular row............................................................................ campestris (Robineau-Desvoidy)

- Postpedicel about 2.5 times as long as broad; scutum with 3 distinct or with 2 indistinct longitudinal vittae, mid vitta usually reduced; presutural acrostichals in 2 rows................................................................. 67

67. Frontal triangle reaching anterior margin of frons.............................................. albicornis Meigen

- Frontal triangle not reaching anterior margin of frons........................................................ 68

68. Scutum with 3 distinct brown longitudinal vittae................................. striolata Hennig & sibirica Hennig

- Scutum without distinct longitudinal vittae, or with 2 indistinct stripes......................... lineatipes (Zetterstedt)

69. Mid femur with a strong anterior preapical seta............................................................. 70

- Mid femur without a strong anterior preapical seta.......................................................... 76

70. At least hind tibiae black.............................................................................. 71

- All tibiae yellow..................................................................................... 73

71. Fore and mid tibiae yellow, hind tibia black; hind tarsi thickened, all tarsomeres about twice as wide as those of mid tarsus ( Fig. 11A )............................................................................ atritibia Ringdahl (part)

- All tibiae predominantly black; hind tarsi not thickened...................................................... 72

72. Scutum with distinct brown narrow longitudinal vittae along dorsocentral, acrostichal setae and posthumeral to intra-alar setae, with brown dust between dorsocentral setae, reaching the apex of scutellum; mid femur with a posteroventral seta in apical third (Figures in Sorokina 2022)....................................................... wrangelensis Sorokina (part)

- Scutum with 3 longitudinal vittae along dorsocentral and acrostichal setae but without brown dust between dorsocentral setae, scutellum grey; mid femur without posteroventral seta in apical third. European Russia..... femoralis (Robineau-Desvoidy)

73. Hind femur with a complete row of posteroventral setae; frontal vitta dull black in frontal view......... perpusilla Meigen

- Hind femur with 0–2 posteroventrals in basal half; frontal vitta grey in frontal view................................ 74

74. Hind femur without posteroventrals in basal half................................................ paludis Tiensuu

- Hind femur with 2 posteroventrals in basal half............................................................ 75

75. Mid femur usually with 1 or 2 anteroventral setae in basal third (long and short setae); mid and hind femora usually yellow in basal third; at least tergites 3–5 with pairs of rounded spots and sometimes a vitta between them........... pumila (Fallén)

- Mid femur with 3 or 4 anteroventral setae in basal half or in basal 2/3; mid and hind femora mostly subshining black; abdomen without or with indistinct pairs of spots...................................................... pilipyga Ringdahl

76. Gena narrower than postpedicel; parafacial narrower than anterior ocellus....................... pygmaea (Zetterstedt)

- Gena distinctly broader than postpedicel; parafacial broader than anterior ocellus.................................. 77

77. All tibiae predominantly black, only basally paler........................................................... 78

- At least fore and mid tibiae yellow....................................................................... 82

78. Scutum with 2–3 brown longitudinal vittae, without brown dust between them; abdomen with distinct brown paired spots... .................................................................................................. 79

- Scutum with brown narrow longitudinal vittae along dorsocentral, acrostichal setae and posthumeral to intra-alar setae; with brown dust between dorsocentral setae, reaching the apex of scutellum; abdomen without spots or with small indistinct brownish marks or spots....................................................................................... 80

79. Scutum with 3 distinct brown longitudinal vittae ( Fig. 4E ); abdomen with large brown paired spots ( Figs 4 F, H ).............................................................................................. octosignata Rondani

- Scutum with 2 distinct brown longitudinal vittae and with traces of a weak median vitta between them ( Fig. 4D ); brown oval spots of abdomen smaller and less well-defined ( Figs 4 B, C )................................. nevadensis Lyneborg

80. Height of gena ca. 1.5–1.7 times width of postpedicel; mid femur with 1 preapical posterior seta, without preapical posteroventral seta; abdomen without spots or marks (Figures in Sorokina 2022)............................... koni Sorokina (part)

- Height of gena ca. 2.0–3.0 times width of postpedicel; mid femur with 1 or 2 preapical posterior setae, with or without preapical posteroventral seta; abdomen usually with dark marks, sometimes indistinct...................................... 81

81. Lower calypter a little longer than upper one; abdomen with brown rounded spots, sometimes indistinct (Figures in Sorokina 2022)........................................................................ wrangelensis Sorokina (part)

- Lower calypter much longer than upper one; abdomen with diffuse brown marks reaching ventral surface of tergites (Figures in Sorokina 2022)................................................................ gorodkovi Sorokina (part)

82. All tibiae completely light yellow ( Fig. 10H ); preapical anterodorsal and posterodorsal setae on hind tibia weak but distinct, each half as long as preapical dorsal seta, this seta a little shorter than median anterodorsal seta; lower calypter a little longer than upper one.................................................................... conflicta Huckett (part)

- Tibiae brownish or partly darkened, not light yellow; preapical anterodorsal and dorsal on hind tibia long and strong, the dorsal about as long as median anterodorsal, without a distinct preapical posterodorsal; lower calypter much longer than upper one. .................................................................................................. 83

83. All trochanters greyish-brown; abdomen with rounded spots, sometimes indistinct; scutum without distinct dorsocentral vittae; hind tibia mostly brownish; postpedicel rounded at apex........................................... verralli Collin

- At least fore and mid trochanters yellow; abdomen usually without spots; scutum with three longitudinal vittae; hind tibia with a dark ring in basal and apical third; postpedicel pointed at apex (Figures in Sorokina 2022).......... tschernovi Sorokina

Figures

FIGURE 1. Coenosia luxia sp. nov. A. Holotype, male, lateral view. B. Holotype, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view. E. Cercal plate, dorsal view. F. Sternite 5. Bare: 1 mm.

FIGURE 4. Coenosia spp. A–D. Coenosia nevaedensis. A. Male habitus, lateral view. B. Female habitus, lateral view. C. Female abdomen, dorsal view. D. Female scutum, dorsal view. E–H. Coenosia octosignata. E. Female scutum, dorsal view. F. Female abdomen, dorsal view. G. Male habitus, lateral view. H. Female habitus, lateral view. I. Coenosia octopunctata, male habitus, lateral view. Bare: 1 mm.

FIGURE 7. Coenosia spp. A, B. Coenosia ciliata. A. Male habitus, lateral view. B. Female habitus, lateral view. C. Coenosia penicillata, male habitus, lateral view. D, E. Coenosia subgracilis. D. Male habitus, lateral view (the holotype of Coenosia altaica Sorokina, 2009). E. Female habitus, lateral view (the paratype of C. altaica Sorokina, 2009). F, G. Coenosia oralis. F. Male habitus, lateral view. G. Female habitus, lateral view; H, I. Coenosia ukokensis. H. Male, holotype, lateral view. I. Female, paratype, lateral view. Bare: 1 mm.

FIGURE 8. Coenosia spp. A, B. Coenosia ozerovi. A. Male habitus, lateral view. B. Female habitus, lateral view. C, D. Coenosia nigrotincta. C. Female habitus, lateral view (Pevek). D. Male habitus, lateral view (Beringovskii). E, F. Coenosia ukokensis. E. Male habitus, lateral view (Magadan). F. Female habitus, lateral view (Magadan). Bare: 1 mm.

FIGURE 9. Coenosia spp. A. Coenosia tigrina, female, hind tibia. B. Coenosia intermedia, female, hind tibiae. C. Coenosia ozerovi, male, fore tarsi. D. Coenosia ambulans, female, head. E. Coenosia kosterini, male, lateral view. F, G. Coenosia alaskensis. F. Female habitus, lateral view. G. Male habitus, lateral view. Bare: 1 mm.

FIGURE 10. Coenosia spp. A, B. Coenosia baikalensis. A. Male habitus, lateral view. B. Female habitus, lateral view. C, D. Coenosia tendipes. C. Female habitus, lateral view. D. Male habitus, lateral view. E, F. Coenosia flaviseta. E. Male, head, lateral view. F. Female habitus, lateral view. G, H. Coenosia conflicta. G. Male habitus, lateral view. H. Female habitus, lateral view. Bare: 1 mm.

FIGURE 11. Coenosia spp. A, B. Coenosia atritibia. A. Male habitus, lateral view. B. Female habitus, lateral view. C, D. Coenosia apukaensis. C. Male habitus, lateral view. D. Female habitus, lateral view. E, F. Coenosia xuei. E. Female habitus, lateral view. F. Male habitus, lateral view. Bare: 1 mm.

FIGURE 12. Coenosia spp. A, B. Coenosia lacustris. A. Male habitus, lateral view. B. Female habitus, lateral view. C, D. Coenosia luteipes. C. Male habitus, lateral view. D. Female habitus, lateral view. E. Coenosia sibirica, male abdomen, ventral view. F. Coenosia lineatipes, male sternite 5, ventral view. G. Coenosia pilipyga, male abdomen, lateral view. H. Coenosia pumila, male abdomen, lateral view. Bare: 1 mm.