Craspedometopon KERTÉSZ, 1909

Rozkošný, R. & Kovac, D., 2007, Palaearctic And Oriental Species Of Craspedometopon Kertész (Diptera, Stratiomyidae), Acta Zoologica Academiae Scientiarum Hungaricae 53 (3), pp. 203-218 : 204-206

Craspedometopon KERTÉSZ, 1909

Craspedometopon KERTÉSZ, 1909: 373 . Type species: Craspedometopon frontale KERTÉSZ, 1909 (by monotypy)

Acanthinoides MATSUMURA, 1916: 367 . Type species: Acanthinoides basalis MATSUMURA, 1916 (= Beris basalis MATSUMURA, 1915 ) (orig. des.).

The genus includes predominantly black and stout flies of small to medium size (5.0–7.0 mm). The head is transverse in both sexes, i.e. much broader than long in dorsal view and higher than long in lateral view. The ocellar tubercle is usually very prominent above the eyes in lateral view. The eyes are bare or microscopically short haired, but finely, relatively long and densely haired in C. orientale sp. n., touching in the males and separated by a parallel-sided frons in the females. The relatively short antennae are inserted below the middle of eyes, each consisting of a cylindrical scape, which is longer than the subconical pedicel, and an onion-like flagellum with an almost terminal arista. The flagellum consists of 7 flagellomeres, forming a subrounded complex, and an arista that is less than twice as long as the antenna ( Fig. 15 ). The face is concave, the labellum of the proboscis large. The two-segmented palpi are moderately long, both segments subequal.

The thorax is black, densely punctate and generally short haired, shining black parts are very limited even on the pleura. The scutum and the scutellum are moderately convex in lateral view, with only a shallow incision between them. The scutellum has four strong and rather short spines and an entire, distinct margin ( Fig. 19 ). The wings are usually infuscated in the basal half, at least the stronger veins in this region are darker than in the distal half. The anterior crossvein is well developed and vein R 2+3 originates well beyond it. Vein R 4 is unusually approximate to R 2+3 so that the costal section between them is much shorter than the section between R 4 and R 5. Vein Cu 2 is markedly bent and the apical angle of cell Cu is thus 90 degrees. The legs are simple, black and yellow, the posterior surface of the fore and mid femur is usually covered with longer erect hairs. The abdomen is wide, rounded or even somewhat broader than long, conspicuously arched.

The male eye facets are markedly enlarged in the upper half, the eyes are contiguous medially for a long distance, leaving only a small and narrow upper frons and a larger and much broader lower frons. The elongate-triangular upper frons is about as broad as the anterior ocellus and abruptly tapered downwards, being barely twice as long as broad. The broadly triangular lower frons is usually covered by dense whitish grey tomentum but it is shining black at the upper angle and in the median groove reaching the base of antennae. The antennal flagellum is, as a rule, smaller than in the female, also the palpi are shorter and more slender. A postocular rim is not visible in dorsal view and in lateral view a somewhat swollen postocular area is developed only in the lower third ( Figs 1 , 16 ). The infuscation of the wings is often more developed in males than in females. The male terminalia are fairly uniform. The epandrium is usually narrowed distally ( Figs 5 , 12 , 20 ). The genital capsule has a narrow ( Figs 7 , 14 ) or broader posteromedial incision ( Fig. 22 ), the gonostyli are of complicated structure, sometimes narrowed distally, with basal lobes and/or inner processes or hooks ( Figs 4 , 11 , 22 ). The aedeagal complex is relatively short, barely extending beyond the proximal margin of the genital capsule, tripartite distally ( Figs 6 , 13 , 21 ).

The female eye facets are not divided and unified in size. The female frons ( Figs 3 , 8 , 18 ) is parallel-sided, usually rather narrow, occupying at most 1/5 of head width, slightly dilated in the upper part. The lower part of the frons is separated by a shining black, low, transverse elevation. The frontal band is margined by a narrow ridge along each eye margin and with large punctures to varying extent, leaving sometimes a differently large longitudinal medial area bare and shining black. The lower frons above the antennae is covered by dense whitish grey tomentum which is divided medially by a subtriangular or oval longitudinal depression. Both frontal tomentose patches are rounded at the upper margin. The postocular rim is only narrow and well visible in dorsal view ( Fig. 8 ) but absent in C. orientale sp. n. ( Fig. 18 ). In lateral view the postocular area in the lower third of the head ( Figs 2 , 17 ) is mostly as swollen as in the male or somewhat larger. The body pile is usually only short and less conspicuous than in the male. The differences between the dark basal half and the pale apical half of the wing is usually less conspicuous in the females. The legs may be more yellow in some species. The female terminalia ( Figs 9–10 ) are elongated, the two-segmented cerci are relatively short, both segments subequal, tergite 9 relatively high. The genital furca is rounded or more oval, with a long and basally dilated proximal middle projection.

Acanthinoides MATSUMURA, 1916 , based on Acanthinoides basalis MATSUMURA, 1916 , from Japan, is a synonym of Craspedometopon . HOLLIS (1963) described a new species, Craspedometopon testacea , which is now considered to be a member of the genus Evaza WALKER ( WOODLEY 2001) . LINDNER (1938, 1941) discussed and illustrated under Acanthinoides basalis a member of another genus. His male should have the bare eyes (cf. his description) but the figured specimen ( LINDNER 1938: fig 133) has long haired eyes and the antennae distinctly differ from Craspedometopon . The figured female originated from Japan (Honshu, Nikko) and might belong to Kolomania PLESKE (see KRIVOSHEINA 1973).

Figures

Figs 8–15. Diagnostic characters of Craspedometopon frontale (8–9, 11–15) and C. orientale sp. n. (10): 8 = female head in frontal view, 9–10 = female terminalia (left) and female genital furca (right), 11 = gonostylus in dorsal view, 12 =dorsal part of male terminalia, 13 = aedeagal complex, 14 = genital capsule, 15 = male antenna. Scales: 0.5 mm (head and antenna), 0.3 mm (female and male

Figs 16–22. Diagnostic characters of Craspedometopon orientale sp. n.: 16 = male head in lateral view, 17–18 = female head in lateral and frontal view, 19 = male scutellum, 20 = dorsal part of male terminalia, 21 = aedeagal complex, 22 = genital capsule.Scales: 0.5 mm (heads), 1.0 mm (scutellum),

Figs 1–7. Diagnostic characters of Craspedometopon basale: 1 = male head in lateral view, 2–3 = female head in lateral and frontal view, 4 = gonostylus in dorsal view, 5 = dorsal part of male terminalia, 6 = aedeagal complex, 7 = genital capsule. Scales:0.5 mm (heads), 0.2 mm (gonostylus), 0.3