Tonkinomeris napoensis, Nguyen & Sierwald & Marek, 2019

Nguyen, Anh D., Sierwald, Petra & Marek, Paul E., 2019, The pill millipedes of Vietnam: a key to genera and descriptions of five new species (Diplopoda: Glomerida: Glomeridae), Raffles Bulletin of Zoology 67, pp. 260-297 : 264-265

Tonkinomeris napoensis , new species

( Figs. 1–4 )

Material examined. Holotype: male (IEBR–Myr 657H) Vietnam, Ha Giang Province, Bac Quang District, Duc Xuan Commune, Na Po Village , forest on limestone, 14–15 April 2014, coll. A. D. Nguyen.

Paratypes: 3 males, 5 females (IEBR–Myr 657P) same data as for holotype .

Etymology. Named after the Na Po Village where type specimens were collected, (feminine adjective).

Diagnosis. The species is differentiated by characters in the genus diagnosis above.

Description. Holotype male (IEBR–Myr 657H). Width of 2 nd tergum 5.0 mm; body length 9.6 mm.

Exoskeleton ( Fig. 1 ): Terga black with median line of yellow triangular spots longitudinally, and two paramedial marbled transverse oval spots ( Fig. 1A, B ). Lateral margins light yellow. Anal shield black with a light brown, triangular spot medially ( Fig. 1D ). Head: Ocelli 5+1, lenses convex, contrasting black against light brown background of head. Tömösváry organs transverse oval, margins slightly concave, ca. 2× wider than long. Antenna clavate apically; antennomere 6 large, ca. 2× longer than wide. Antennal tip with four large, apical sensory cones. Tergites: Collum semi-circular, with a very large marbled yellow-brown spot medially ( Fig. 1C ). Second tergum (=thoracic shield, tho) with a narrow hyposchism (hs), reaching to caudal margin; dorsal surface with 6–7 distinct transverse striae, all crossing dorsum. Anal shield strongly rounded, emarginated mediocaudally ( Fig. 1D ). Other tergites each with 3–4 superficial striae. Legs: Leg-pair 17 strongly reduced, 3-segmented: coxa with a regular outer lobe, and a setiferous tubercle distomesally; podomere 1 with a small distomesal setiferous knob. Legpair 18 ( Figs. 2A, C , 3A, B ), robust, stout, 4-segmented: coxa with syncoxial notch (arrow); podomere 2 large, with sparsely setose edge (ed) mesally; podomere 3 with a short, stout, sparsely setose tubercle (stu) on posterior side ( Fig. 2C ); podomere 4 with apical seta.

Telopods ( Figs. 2D, E , 3D, E , 4 ): Telopods with a rounded trapeziform, central syncoxial lobe (syl) with two setose horns (syh) paramedially, each projected mesoventrad above syncoxial lobe. Prefemur broadened basally, carrying a short, stout tuberculiform trichostele (pret). Femur with a straight, distal process (dfp), without femoral trichosteles. Tibia with a large, densely microgranulated, lamelliform process (tp). Tarsus rounded apically, and not sigmoid, carrying a seta; mesal side sparsely setose.

Variation. Width of 2nd tergum 3.8–5.0 mm (males), 5.2–6.5 mm (females); body length 5.3–9.6 mm (males), 6.9–9.6 mm (females). A particular big female 12.6 mm long, width 10 mm at tergum 2.

Remarks. Tonkinomeris napoensis , new species is only known from its type locality.

Figures

Fig. 1. Tonkinomeris napoensis new species, holotype (IEBR–Myr 657H), Na Po village (Ha Giang Prov.). A, whole body, dorsal view; B, lateral view; C, ventral view. D, anal shield, posterior view. Scale bar = 1 mm.

Fig. 2. Tonkinomeris napoensis new species, holotype (IEBR–Myr 657H), Na Po village (Ha Giang Prov.). A, legs 18, anterior view; B, posterior view; C, lateral view. D, telopods, anterior view; E, posterior view. Scale bar = 1 mm.

Fig. 3. Tonkinomeris napoensis new species, holotype (IEBR–Myr 657H), Na Po village (Ha Giang Prov.). A, legs 18, anterior view; B, right leg 18, anterior view; C, telopods, anterior view; D, posterior view. Scale bar = 1 mm (A, C); 0.5 mm (B); 2 mm (D).

Fig. 4: Tonkinomeris napoensis new species, holotype (IEBR–Myr 657H), Na Po village (Ha Giang Prov.). A,telopods: syncoxial lobe and horns, posterior view; B, anterior view; C, left telopod, distal part, anterior view; D, basal part, anterior view. Scale bar = 0.3 mm (A, B); 0.5 mm (C, D).