Exocelina pseudoedeltraudae Shaverdo & Balke

Shaverdo, Helena, Sagata, Katayo, Panjaitan, Rawati, Menufandu, Herlina & Balke, Michael, 2014, Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 468, pp. 1-83 : 21-23

Taxon classification Animalia Coleoptera Dytiscidae

15. Exocelina pseudoedeltraudae Shaverdo & Balke sp. n. Figs 9, 38; 4 A–F, 30 in Shaverdo et al. (2012)

Exocelina undescribed sp. MB1288: Toussaint et al. 2014: Supplementary figs 1-4, Tab. 2.

Type locality.

Papua New Guinea: Hela Province, Koroba, 05°41.85'S; 142°43.836'E.

Type material.

Holotype: male "Papua New Guinea: Southern Highlands, Koroba, 1600m, 15.v.1994, 05.41.854S 142.43.836E, Balke (PNG 66)", "Paratypus Exocelina edeltraudae sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012" (ZSM). Paratypes: 5 males, 1 female with the same labels as the holotype (NHMW, ZSM). 3 males "PAPUA N.G.: 6.-9.5.1998 Southern Highl. Prov. Tari-Koroba, Hedemari [Hedamali] 1700-1900 m, leg. Riedel", "Paratypus Exocelina edeltraudae sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012" (NHMW). 1 male, 3 females "Papua New Guinea: Southern Highlands, Tari Komo road, 10km N Hides Gas, 1700m, 13.v.1994, Balke (PNG 61)", "Paratypus Exocelina edeltraudae sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012", the male additionally with a green label "DNA M.Balke 1288" (ZSM). 5 males, 8 females "Papua New Guinea: Southern Highlands, Tari to Koroba, 1600m, 15.v.1994, 05.46.500S 142.50.000E, Balke (PNG 65)", "Paratypus Exocelina edeltraudae sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012" (NARI, NHMW, ZSM). 1 female "Papua New Guinea: Southern Highlands, Tari to Koroba, 1600m, 15.v.1994, 05.46.500S 142.50.000E, Balke (PNG 65)" (ZSM).

Diagnosis.

Beetle medium-sized, piceous, submatt; pronotum with distinct lateral bead; male ventrite 6 slightly to distinctly concave apically; male antennomeres 3-5 distinctly enlarged, almost equal in size and shape, antennomeres 6-8 enlarged; male protarsomere 4 with large, slender, evidently curved anterolateral hook-like seta; median lobe with very strong submedian constriction and proximal part very broad in ventral view, apex of median lobe pointed and strongly curved downwards in lateral; paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, long, thin setae. The species is very similar to Exocelina edeltraudae Shaverdo, Hendrich & Balke, 2012, from which differs with less shinier dorsal surface, due to stronger punctation and microreticulation, with larger and more rounded male antennomeres 3-5 (for male antennomeres 3 and 4, ratio width/length:> 1.0) and apex of median lobe narrower in lateral view.

Description.

Size and shape: Beetle medium-sized (TL-H 3.45-4.0 mm, TL 3.85-4.45 mm, MW 1.8-2.2 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsally piceous, with dark brown anterior margin of head and narrowly pronotal sides; head appendages and legs reddish to reddish brown, legs distally darker (Fig. 38). Teneral specimens paler.

Surface sculpture: Head with dense, coarse punctation (spaces between punctures 1-3 times size of punctures), especially on vertex. Pronotum with punctation finer, sparser, and more evenly distributed than on head. Elytra with punctation finer, sparser than on pronotum. Pronotum and elytra with strongly impressed microreticulation, dorsal surface submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal figs with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.

Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded and smooth anteriorly, with small anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded, or slightly truncate, or concave apically.

Male: Antennomeres 3-5 distinctly enlarged, almost equal in size, antennomeres 6-8 enlarged (Fig. 9A), antennomeres 3-7 rugose ventrally. Protarsomere 4 with large, slender, evidently curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 15 and posterior row of 5 short setae (Fig. 9B). Abdominal ventrite 6 with 8-13 lateral striae on each side, slightly to distinctly concave apically (Fig. 9C). Median lobe with very strong submedian constriction and proximal part very broad in ventral view, apex of median lobe pointed and strongly curved downwards in lateral view (Fig. 9D, E). Paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, long, thin setae (Fig. 9F).

Holotype: TL-H 4.0 mm, TL 4.45 mm, MW 2.2 mm.

Female: Antennae simple, abdominal ventrite 6 broadly rounded or slightly truncate apically, without striae.

Distribution.

Papua New Guinea. The species is known only from Hela Province (Fig. 53).

Etymology.

In an earlier work ( Shaverdo et al. 2012), this species was mistaken for Exocelina edeltraudae . The name is a noun in the nominative singular standing in apposition.