Lycopodium clavatum ssp. clavatum

Øllgaard, Benjamin, 2020, Lycopodiaceae in Colombia: Subfamilies Lycopodioideae and Lycopodielloideae, Phytotaxa 433 (3), pp. 195-224 : 198

Lycopodium clavatum ssp. clavatum — Fig. 2C .

Lycopodium aristatum Willdenow (1810: 17) .— L. clavatum L. var. aristatum (Willd.) Spring (1838: 173) . Type:— VENEZUELA: Silla de Caracas, Humboldt & Bonpland s. n. (B-Willd. 19351 holotype).

Lycopodium piliferum Raddi (1825: 79) .— L. clavatum L. var. raddianum Spring (1838: 172) .— L. clavatum L. var. piliferum (Raddi) Nessel (1927: 435) . Type:— BRAZIL: Mandiocca, G. Raddi s.n. (lecto- PI; isolecto- BM, FI-3 sheets, K, P, PI-2 sheets).

Lycopodium serpens Presl (1825: 81) , non Poiret (1814).— Lycopodium preslii Grev. & Hook. (1831: 377) .— L. clavatum L. var. preslianum Spring (1842: 90) . Type:— ECUADOR: Guayaquil, Haenke s. n. ( PRC holotype).

Lycopodium trichophyllum Desvaux (1827: 184) , [as trychophyllum].— L. clavatum L. var. desvauxianum Spring (1838: 173) .— L. aristatum Willd. var. desvauxianum Spring in Martius (1840: 114).— L. trichiatum Bory var. desvauxianum Spring (1842:92) .— L. clavatum L. var. trichophyllum (Desv.) Nessel (1927:436) . Lectotype:— BRAZIL: »Habitat in Brasilia, no. 41 in herb. de A. N. Desvaux, Donnée par Mme Vve Lavallée en 1895« (P, designated by Øllgaard (1988: 125).

Lycopodium eriostachys Fée (1869: 224) .— L. clavatum L. var. eriostachys (Fée) Nessel (1927: 435) . Type:— BRAZIL: Brasilia fluminensi, Serra os Orgaos, Glaziou 1788 (BR!, C!, P!, RB!).

Lycopodium clavatum L. var. minarum Christ (1900: 41) . Types:— BRAZIL: Minas Geraes: Serra do Caparaó, 1800 m, Schwacke 12001 (P, lectotype, designated by Øllgaard & Windisch (2014: 300)); Serra do Picú, Schenk 1522 (P).

Lycopodium clavatum L. var. equisetoides Nessel (1927: 438) . Types:— BRAZIL: Minas Geraes: Wawra (W, not seen) ; São Paulo: Grossmann 36, 1904 (not located); Paraná: Dusén s. n. 18/11/898 (S, not seen, illustrated in Nessel (1955: fig. 103), presumably showing a duplicate specimen from S now in BONN-Herb. Nessel).

Habitats: Clearings and exposed habitats in montane forest, alt. 1000–3500 m. Terrestrial and rupestral, trailing or scrambling in open and usually humid places like road banks, roadsides, rock crevices, trail margins, often in pioneer habitats.

Antioquia: Medellín–Puerto Triunfo, 75°10’W 6°10’N, 2090 m, Juncosa 965 (AAU). Medellín– Santa Rosa road, 17 km S of Santa Rosa, 2320 m, Luteyn et al. 7026 (AAU). Yarumal–Valdivia road 10 km N of Yarumal, 2135 m, Luteyn et al. 7067 (AAU). Mun. Urrao, Vereda El Chuscal, Jaiperá, El Morro, 76°46’W 6°27’N, 2600–3200 m, Callejas et al. 7877 (NY). Mun. Belmira, Old road Belmira–Sta. Rosa, 2900–3000 m, Arbeláez 144 (AAU). Caldas: Manizales–Neira road, 3 km from Manizales, 2030 m, Santa & Hoyos 675 (AAU). Nevado del Ruiz, road Manizales–Hotel Termales, 3080 m, Santa & Hoyos 678 (AAU). Cord. Occidental, W slope, La Selva, 1600–1900 m, Sneidern 5444 (AAU, M).

Caquetá: Mun. Florencia, road Florencia–Neiva, Quebrada La Portada, 1200 m, Santa 1232 (AAU). Chocó: Mun. San José del Palmar, Cerro del Torrá, E slope, 2500 m, Silverstone-Sopkin 4571 (AAU). Mun. San José del Palmar, Cerro del Torrá, summit, 2700–2800 m, Silverstone-Sopkin 4654 (AAU). Mun. San José del Palmar, Cerro del Torrá, E slope, 2500 m, Silverstone-Sopkin 4286 (AAU). Cundinamarca: Mun. Bojacá, San Antonio trail, La Merced, near Mosquera–Tena Road, 2500–2700 m, Lozano-C. & Torres-R. 75 (AAU). Mun. Albán. road Bogotá –Sasaima, vicinity of La Frontera, 2000 m, Acosta Arteaga 360 (AAU). Bogotá –Usme road, Páramo de Chisacá, km 28–33, 3375–3450 m, Luteyn et al. 7746 (AAU). Huila: Paramillo Arrabal, Finca La Candelaria, W of Volcán Puracé, near Santa Leticia , 76°15’W 2°16’N, 2380 m, Gentry et al. 54035 (AAU). Magdalena: Sierra Nevada de Santa Marta , road to Cuchilla de San Lorenzo, above Estación de Telecom, 1840 m, Santa & Escobar 635 (AAU). Sierra Nevada de Santa Marta , crest of San Lorenzo, 2750 m, Santa & Escobar 663 (AAU). Nariño: Pasto, Jameson 410 (G). Ricaurte, La Planada, 5 km S of Chucunés, 1850 m, Luteyn 6833 (AAU). Putumayo: Páramo de Tambillo, NE of Valle de Sibundoy, 2700–2800 m, Schultes & Smith 3114 (MO, NY). Santander: Subpáramo and forest NW of Arabuco, Cerro Berlín, Río Chuqueque drainage, 73°30’W 5°40’N, 2825 m, Cleef et al. 3492 (AAU). Valle: Ansermanuevo– San José del Palmar road, km 60–62, 1870–1950 m, Luteyn et al. 7271 (AAU). Mun. El Cairo, Las Amarillas, 1 hour’s drive from El Cairo Valle near base of Cerro del Inglés, Serranía de Los Paraguas, E slope of Cord. Occidental, 2070–2125 m, Silverstone-Sopkin 3751 (AAU).

Figures

FIGURE 2. A: Diphasiastrum thyoides (Ecuador: Holm-Nielsen et al. 4818, AAU), branchlets seen from above (left) and below (right). B: Diphasium jussiaei (Ecuador: Holm-Nielsen et al. 4670, AAU) branchlets seen from (left) and below (right). C: Lycopodium clavatum ssp. clavatum (Ecuador. Harling 4091, S). C1: Branchlet leaf. C2: Sporophyll. D: Lycopodium clavatum ssp. contiguum (Ecuador: Cazalet & Pennington 5409, US). D1: Sporophylls.