Lacon ganglbaueri ( Schwarz, 1894 )

Kundrata, Robin, Mertlik, Josef & Németh, Tamás, 2019, Unexpected diversity of Lacon Laporte, 1838 (Coleoptera: Elateridae: Agrypninae) in the Levant: revised species concepts, new species, and an identification key, Zootaxa 4679 (3), pp. 401-449 : 415-417

Lacon ganglbaueri ( Schwarz, 1894)

( Figs 52–64 , 188 , 194 )

Adelocera ganglbaueri Schwarz, 1894: 145 .

Adelocera ganglbauri , Lacon ganglbauri: Hayek (1973: 65) ; unavailable name, incorrect subsequent spelling not in prevailing usage ( ICZN 1999, Art. 33.3).

Lacon ganglbaueri: Fleutiaux (1926: 93) ; see Hayek (1973: 14, 65).

Type material. Lectotype, male, „Appl Beirut [ Lebanon] 1878, coll. Schwarz, Syntypus, DEI Eberswalde, Ganglbaueri Schw., Lectotypus Adelocera ganglbaueri Sch. , ♂, des. Platia 1997, Lacon ganglbaueri (Scw) det. Platia 1997“ ( SDEI). Paralectotype, female, „Appl Beirut 1878, B. M., Ad. Ganglbaueri det. Schwarz, Als Typus festgestellt XI.63, Schönmann, Typus, Lacon ganglbaueri bona sp., det. C. Wurst 1997“ ( NHMW).

Type locality. Lebanon: Beirut .

Other material examined. Lebanon, 2 males, „Lebanon: Ajaltun, Beirut, 3000 feet, H. St. J. Philby [leg.]; Date: 27.V.1956, B. M. 1958-702, Lacon bruleriei ??, ♂, det. Preiss, 2003 “ ( BMNH); 1 male, „ Beirut, Sharp Coll., 1905-313., Lacon ganglbaueri ??, ♂, det. Preiss 2003 “ ( BMNH); 1 male, „Beyrout, Sharp Coll., 1905-313., lithophila “ ( BMNH); 1 male, 1 female, „Lebanon, Mount Leb. Range, Arsoun, Pinus forest, from Tomicus pheromone trap, 33°51‘8.03“N, 35°41‘44.36“E, X.2016, N. Nemer [leg.] (No. 42/d)” ( HNHM); 3 males, “n. LIBANO caza Bcharre, Wadi Kadicha m. 900, giu 2008, leg. A. Kairouz, Lacon ganglbaueri (Schwz.) det. Platia 2009 [one specimen with „det. Platia“ only]” ( PCGP); 1 female, “Lebanon, Ref.17Ca, IIE 23988, Ain Saade 17Ca, Lebanon, Pinus pinea ecosyst., 2/07/1997, Routier/Nemer, Lacon sp. near drusa (Mars.) det. R. G. Booth, 1999, Elateridae , Lacon sp. (Marseul) det. R. G. Booth ( IIE), Lacon ganglbaueri (Schw.) det. Platia 2015 “ ( HNHM).

Comparative remarks. This species shares the general habitus, male antenne surpassing pronotal posterior angles, and the sclerites of bursa copulatrix in females with L. candezei , L. carmelensis (female unknown), and L. solai . The most similar species to L. ganglbaueri is L. candezei . These species share also the body size, pubescence with a combination of greyish to reddish-brown/dark brown and white setae, and a median lobe of aedeagus distinctly shorter than parameres ( Figs 36–46 , 52–59 ). For the discussion about the differences between these species see the „comparative remarks“ under L. candezei . Lacon solai from Turkey differs from L. ganglbaueri in having usually larger body (10.0– 14.5 mm long), and a median lobe of aedeagus longer, reaching apical lobes of parameres ( Figs 52–55, 59 , 70–73, 77 ). Lacon carmelensis from Israel differs from L. ganglbaueri in having a darker body, body pubescence with mostly dark brown and some white setae, and the aedeagus without phallobase stouter, less than four times longer than wide (more than 4.2 times longer than wide in L. ganglbaueri ), and a median lobe of aedeagus relatively longer, not reaching parameral lateral tooth of only about 0.5 length of the apical parameral lobe (of about the length of the apical lobe or more in L. ganglbaueri ) ( Figs 52–55, 59 , 65, 66, 69 ).

Diagnostic characters. Male ( Figs 52–54, 56, 58, 59 ). Body 8.3–9.8 mm long and 2.3–2.6 mm wide (lectotype: 8.5 mm long, 2.3 mm wide). Body moderately shiny, reddish brown to dark brown, with antennae, pronotal posterior angles and legs ferruginous; covered with mostly greyish to reddish brown/dark brown setae (some of them tricolorous: greyish, reddish brown, and white) and several white setae, all moderately long and thick.Antenna relatively long, surpassing posterior angles of pronotum usually by about 1.0–1.5 antennomeres. Antennomere III 1.15–1.25 times as long as antennomere IV, median antennomeres rather widely serrate, about as long as wide. Pronotum 1.1–1.2 times as long as wide, with posterior angles slightly divergent, disk densely covered with distinct, large, deep punctures, separated by about 0.5–1 times as diameter of puncture; surface between punctures flat, shiny. Scutellar shield about 1.5 times as long as wide, with anterior margin slightly concave. Elytra 2.3–2.4 as long as wide, and 2.4–2.6 times as long as pronotum, with punctures of two sizes; smaller punctures of about 2/3 of size of larger punctures. Aedeagus (without phallobase) slender, more than 4.2 times longer than wide; median lobe short, not reaching lateral hook of paramere of about the length of the apical parameral lobe; parameres subparallel-sided along most of their length, apically with inner sides distinctly diverging; apical parameral lobe elongate, narrow, with outer margin more or less straight, apically subacute [lectotype specimen already dissected; basal part of aedeagus including phallobase missing ( Fig. 59 )].

Female ( Figs 55, 57, 60–64 ). Body 9.2–9.5 mm long and 2.5–2.7 mm wide. Body usually more robust than in male. Antenna shorter, not reaching 2/3 of pronotal length. Antennomere III 1.20–1.35 times as long as antennomere IV; median antennomeres of similar shape as in male. Pronotum relatively shorter and wider, sides widely rounded, 1.05–1.10 times as long as wide, sides equally rounded, posterior angles slightly convergent. Elytra combined 2.1–2.2 times as long as wide, about 2.1 times as long as pronotum. Pregenital segments, ovipositor and sclerites of bursa copulatrix as in Figs 60–64 , respectively.

Immature stages unknown.

Distribution. Lebanon ( Fig. 194 ). For the habitat photograph see Fig. 188 .

Literature. Schwarz (1894: 145): original description; Schwarz (1906: 11): catalogue; Schenkling (1925: 10): catalogue; Fleutiaux (1926: 93): catalogue, nomenclatural note; Hayek (1973: 65): revision; Cate (2007: 101): catalogue; Németh (2019: 129): catalogue.

Figures

FIGURE 194. Distribution of Lacon spp. in the mainland Levant. Dots representing L. candezei in Lebanon and Syria are only approximate since there are no exact data for these countries.

FIGURES 52–64. Morphology of Lacon ganglbaueri. Habitus: 52–53, lectotype, male, dorsal and lateral view, respectively; 54, male from Lebanon, dorsal view; 55, female from Lebanon, dorsal view. Pronotum, dorsal view: 56, male lectotype; 57, female paralectotype. Lectotype, male: 58, detail of elytral surface; 59, aedeagus. Female paralectotype, pregenital segments and genitalia: 60, tergite VIII; 61, sternite VIII; 62, ovipositor; 63, large sclerite of bursa copulatrix. 64, female from Lebanon, large sclerite of bursa copulatrix. Scale bars = 2.0 mm (Figs 52–55), 1.0 mm (Figs 56–57), 0.5 mm (Figs 59–62), 0.2 mm (Figs 63–64); Fig. 58 not to scale.

FIGURES 188–193. Habitat and live specimens of Lacon spp. 188, Pinus pinea ecosystem at Arsoun, Lebanon, habitat of L. ganglbaueri and L. punctatus. 189, L. punctatus. 190, Cedrus lebani ecosystem at Tannourine Cedar Forest Nature Reserve, Lebanon, habitat of L. drusus. 191, L. drusus. 192, macchia at Erateini, Greece, habitat of L. graecus. 193, L. graecus.

FIGURES 36–51. Morphology of Lacon candezei. Habitus: 36–37, lectotype, male, dorsal and lateral view, respectively; 38, male from Jordan, dorsal view; 39, female from Jordan, dorsal view. Pronotum, dorsal view: 40, lectotype, male; 41, male from Jordan; 42, female from Jordan. 43, lectotype, male, detail of elytral surface. Male genitalia: 44, lectotype; 45, specimen from Jordan; 46, specimen from Israel. Female pregenital segments and genitalia of specimen from Jordan: 47, tergite VIII; 48, sternite VIII; 49, ovipositor; 50, sclerites of bursa copulatrix. 51, female from Israel, sclerites of bursa copulatrix. Scale bars = 2.0 mm (Figs 36–39), 1.0 mm (Figs 40–42, 48–49), 1.5 mm (Figs 44–47), 0.2 mm (Figs 50–51); Fig. 43 not to scale.

FIGURES 70–79. Morphology of Lacon solai. Habitus: 70–71, holotype, male, dorsal and lateral view, respectively; 72–73, female, dorsal and lateral view, respectively. Pronotum, dorsal view: 74, holotype, male; 75, female. 76, holotype, male, detail of elytral surface. 77, holotype, male genitalia. 78, female, ovipositor. 79, female, sclerites of bursa copulatrix. Scale bars = 2.0 mm (Figs 70–73, 78), 1.0 mm (Figs 74–75), 0.5 mm (Figs 77, 79); Fig. 76 not to scale.

FIGURES 65–69. Morphology of Lacon carmelensis, holotype, male. 65, habitus, dorsal view; 66, habitus, lateral view; 67, pronotum, dorsal view; 68, detail of elytral surface; 69, aedeagus. Scale bars = 2.0 mm (Figs 65–66), 1.0 mm (Fig. 67), 0.5 mm (Fig. 69); Fig. 68 not to scale.