Pachyptera DC., Prodr. 9: 175. 1845

Francisco, Jessica Nayara Carvalho & Lohmann, Lucia G., 2018, Taxonomic revision of Pachyptera (Bignonieae, Bignoniaceae), PhytoKeys 92, pp. 89-131 : 97-98

Pachyptera DC., Prodr. 9: 175. 1845

Pachyptera DC., Prodr. 9: 175. 1845. Type: Pachyptera foveolata DC. (lectotype, designated by Sandwith [1932: 84]) [= Pachyptera kerere (Aubl.) Sandwith]

Sererea Raf., Sylva Tellur. 107. 1838. Type: Sererea heterophylla (Willd.) Raf., Sylva Tellur. 107. 1838. nom. illeg. superfl. [= Pachyptera kerere (Aubl.) Sandwith]

Leucocalantha Barb. Rodr., Vellosia, ed. 2. 1: 46, tab. 7. 1891. Type: Leucocalantha aromatica Barb. Rodr., Vellosia. ed. 2. 1: 47, tab. 7. 1891.

Description.

Liana; stems with four phloem wedges in cross-section, solid (hollow in some specimens of P. aromatica ), cylindrical to tetragonal, striated, with lenticels (without in P. aromatica ), with interpetiolar extrafloral nectaries, with a continuous (discontinuous) and transversal interpetiolar ridge, with a papery peeling bark, lepidote, puberulous becoming glabrescent with age; prophylls of the axillary buds 3(-5) seriated (a single series visible to the naked eye in some specimens of P. aromatica ), flattened and ensiform (triangular and minute in P. aromatica ). Leaves 3-2-foliolate with the terminal leaflet replaced by a trifid tendril; blades discolorous (concolor), chartaceous to coriaceous, usually asymmetric (symetric in some specimens of P. aromatica ), apex mucronulate, glabrous to puberulous, with simple trichomes covering veins (throughout surface), lepidote, with patelliform trichomes throughout the lamina, venation pinnate, secondary venation brochidromous, tertiary venation percurrent, margin entire, flat or sub-revolute; petioles striated, apices articulated, glabrous to puberulous, lepidote, patelliform glands distributed at petiole apices; petiolules with unequal lengths, striated, apices not-pulvinated (pulvinated in P. aromatica ), puberulous, lepidote, lateral petiolules shorter than the apical ones. Inflorescence axillary or terminal, a few-flowered raceme, congested (lax in P. aromatica ); axis puberulous, lepidote, patelliform glands grouped at the axis; pedicel puberulous, lepidote; bracts and bracteoles caducous, scarcely evident, puberulous, lepidote. Calyx tubular (cupular in P. erythraea ), coriaceous, smooth, glabrous internally, puberulous externally, lepidote, patelliform glands grouped at the upper portion. Corolla white to cream (orange to red in P. erythraea and light pink to pale purple in P. incarnata ), with yellow nectar guides, infundibuliform, (hypocrateriform in P. aromatica and tubular-campanulate in P. erythraea ), straight, dorso-ventrally compressed (not compressed in P. aromatica ), membranaceous, tube puberulous externally, lepidote, glabrous internally, but villous at the region of insertion of stamens and staminode (glabrous in P. aromatica ); lobes imbricate, with a pair of patelliform glands arranged in lines externally, lepidote internally. Androecium didynamous, included in two heights, with one staminode, glabrous (puberulous in P. erythraea ); anthers white, becoming darkish with age, included (sub-exserted in P. erythraea ), villous (glabrous in P. aromatica ), basifixed, connective thick, round (acute in P. aromatica ), with thecae divergent, curved forward (straight in P. aromatica ); pollen 3 colpate, microreticulate (4 colpate, psilate-foveolate-microreticulate in P. aromatica ). Gynoecium glabrous; ovary cylindrical, not-sulcate (bisulcate in P. erythraea ), smooth, pubescent, (densely lepidote in P. erythraea and P. incarnata ); ovules arranged in two series per locule, placentation axial; stigma glabrous; nectar disc well developed, glabrous. Capsule linear, flattened (fusiform, inflated in P. kerere ), coriaceous to woody, smooth, puberulous, lepidote, with patelliform glandular trichomes throughout, in higher densities at the margins of valves, without lenticels, each valve with an inconspicuous longitudinal midline (conspicuous and raised in P. kerere ), calyx caducous; seeds oblong, thin, not-corky (irregulary circular, obcordate, thick and corky in P. kerere ), chartaceous to coriaceous, glabrous, smooth, striated, winged (wingless in P. kerere ), with membranaceous (chartaceous) and hyaline wings.

Nomenclatural note.

Sererea Raf. was described to accommodate one species, Sererea heterophyla Raf., using a wrong spelling. However, Sererea heterophylla (Willd.) Raf. was actually based on Bignonia heterophylla Willd, a superfluous name for Bignonia kerere Aubl.

Number of species, distribution and habitat.

Pachyptera comprises five species found in wet and flooded forest vegetation from Belize to Bolívia and Brazil.

Key to species of Pachyptera