Incasarcus argenteus Kury & Maury, 1998

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

3.31. Incasarcus argenteus Kury & Maury, 1998

Figs 4B , 9A, B , 13E, F , 30

Incasarcus argenteus Kury & Maury, 1998: 155 (desc.), 159 (key), figs. 32 (male dorsal habitus, chelicerae, pedipalpus), 33 (penis dorsal view), 34 (penis lateral view), 35 (male lateral habitus, chelicera, pedipalpus), 36 (female dorsal habitus, chelicerae, pedipalpus), 37 (male trochanter-tibia IV); Kury 2003: 144 (cat.); Benavides et al. 2021: 651(cit) fig.1 (cladogram).

Redescription.

MALE: Measurements (n =3) DSW: 5.5-6.1 (6.1); DSL: 6.1-7.4 (7.4); CL: 2.5-3.1 (3.1). FIVL: 9.7-10.8 (10.8). ChL: 3.4-5.2 (5.2). Coloration (in ethanol): Carapace (more accentuated behind and next to ocularium), area I and lateral margins of dorsal scutum, free tergites I-II, coxa IV (dorsal and ventral surfaces) and free sternites white-silver. Remaining mesotergum, pedipalpus and legs dark brown. Chelicerae yellowish brown. Dorsum: (Fig. 4B ) Anterior margin of dorsal scutum granulate. Ocularium with well-defined median depression; granulate. Carapace entirely covered by granules. Areas I-IV granulate, densely distributed in areas I-II and very sparsely distributed in areas III-IV; I with a pair of median tubercles; II-IV a pair of small tubercles near the lateral regions (absent in some specimens); III with a median pair of small or large spines, directed upwards. Lateral margins of dorsal scutum entirely covered by granules (irregularly distributed) except close to areas I-IV. Posterior margin of dorsal scutum predominantly smooth, with a row of granules. Free tergites I-III with irregular row of tubercles of different sizes, the largest and acuminate in the median portion. Chelicerae: (Fig. 4B ) Swollen. Segment I with four small granules. Segment II predominantly smooth, with four teeth. Segment III with three teeth. Pedipalpus : Femur slightly granulate dorsally. Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 7-8 large setiferous tubercles, divided into two groups, one basal with two tubercles and the remaining occupying the median portion of the segment; apical portion smooth; one large proapical spine. Patella with a proapical tubercle. Tibial: prolateral iiiIii, retrolateral IiIi. Tarsus: prolateral iiIiIi / iiiIiiIi, retrolateral iIiIii. Venter: Coxa I with a median row of seven setiferous tubercles. Coxae II-IV covered by setiferous granules. Rows of four small tubercles between coxae II-III and seven between the coxae III-IV. Stigmatic area slightly granulate. Free sternites I-III each with row of granules. Anal operculum granulate. Legs: (Figs 4B , 9A, B ) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with few scattered granules. Trochanters I-III unarmed and granulate. Trochanter IV with a retroapical tubercle. Femora I-III unarmed and with scattered small granules. Femur IV covered with small granules; a retroventral row of 35-37 tubercles of equal size arranged along the entire length of article; a proventral row of 31-32 acuminate tubercles, the largest on the median portion; a prolateral row with 14 tubercles along the basal ⅓. Patellae I-III unarmed. Patella IV with a retrodorsal apical large tapered apophysis; a smaller apical prodorsal apophysis; a retroventral row of five acuminate tubercles and a proventral row of three tubercles. Tibiae I-IV unarmed and granulate. Tarsal segmentation: (n =3) 10-11 (10), 17-19 (19), 10-11 (11), 12-13 (13). Penis: (Fig. 13E, F ) Truncus swollen apically. VP rectangular with straight distal margin; straight in lateral view. MS C1-C3(C4) subapical long and apically curved; MS sub basal A1-A2 short; MS D1 very short, more dorsally placed (near MS C). Lateral sacs with long base, short length, apically blunt; with long T3-like microsetae. Stylus with apex dorsoventrally slightly inflated; with small apical projections. Promontory convex. - FEMALE: Measurements (n =5) DSW: 4.3-5; DSL: 4.9-6.0; CL:1.6-2.3. FIVL: 9.7-10.9. ChL: 1.4-1.9. Tibia: prolateral iiIi / iIiIi. Tarsus: prolateral iiiIiI / iiiIiIi, retrolateral iIiIi. Chelicerae not swollen as in male. Femur-patella IV unarmed. Areas I-II and IV of dorsal scutum unarmed. White-silver only laterally behind ocularium and on area I of dorsal scutum. Tarsal segmentation: (n =5) 9, 16-18, 10-11, 10-12.

Diagnosis.

It differs from other species of the genus by silver-white patches on carapace, area I and lateral margins of dorsal scutum; femur IV with two rows of acuminate large tubercles, a retroventral one with 35-37 tubercles and a proventral one with 31-32 tubercles (Fig. 9A, B ).

Distribution.

(Fig. 30 ) PERU. Cusco. La Convención and Urubamba provinces.

Material examined.

Type material: Holotype ♂, ' PERU, Cusco, Urubamba province, Ollantaytambo district, Abra de Malaga , Cancayoc , 3,000 m a.s.l., 13°16′S 72°16′W, 27/VIII/1995, J. Ochoa leg. (MACN 9549) - Paratype 1 ♀, ‘ditto’ (MACN 9550). Additional material: 2 ♂, 4 ♀, ' PERU, Cusco, La Convención province, Carrizales , 3,250m a.s.l., 13/IV/2014, R. Cruz, S. Bejar & M. Serrano leg. (MZSP 76552) .

Figures

Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury & Maury, 1998; C I. dianae Kury & Maury, 1998; D I. ochoai Kury & Maury, 1998; E I. pictus Kury & Maury, 1998; F I. viracocha Kury & Maury, 1998; Legend bars = 1 mm.

Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury & Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury & Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury & Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury & Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury & Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm.

Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury & Maury, 1998; G - H Incasarcus dianae Kury & Maury, 1998; I - J: Incasarcus pictus Kury & Maury, 1998; Legend bars = 0.1 mm.

Figure 30. Distribution of Incasarcus in Cusco, Peru.