Socalchemmis, PLATNICK & UBICK, 2001

PLATNICK, NORMAN I. & UBICK, DARRELL, 2001, A Revision of the North American Spiders of the New Genus Socalchemmis (Araneae, Tengellidae), American Museum Novitates 3339, pp. 1-26 : 2-3

SOCALCHEMMIS , NEW GENUS

TYPE SPECIES: Anachemmis dolichopus Chamberlin.

ETYMOLOGY: The generic name is a contraction of southern Californian Chemmis , and is masculine in gender .

DIAGNOSIS: Males can be distinguished from those of the other North American tengellid genera by having both a bipartite embolus (fig. 2) and a bipartite retrolateral tibial apophysis (fig. 3); females have a large epigynal septum (figs. 4, 34) and a flatter, less three­dimensional epigynum than do those of the other genera.

DESCRIPTION: Medium to large spiders, total length of males 7.1–13.2, of females 7.5– 15.2. Carapace oval, widest at rear of coxae II, abruptly narrowed at level of palpi to less than half of maximum width; thoracic groove long, longitudinal, very deep; surface coated with short recumbent and fewer, longer, erect dark setae, erect setae most numerous in ocular area; eight eyes in two rows; from above, both eye rows slightly recurved; from front, anterior row recurved, posterior row slightly procurved; anterior median eyes round, smallest; other eyes oval, subequal, with canoe­shaped tapeta; anterior median eyes separated by roughly their diameter, slightly closer to anterior laterals; posterior medians separated by roughly their diameter, farther from posterior laterals; lateral eyes of each side separated by less than their diameter; median ocular quadrangle wider in back than in front, wider in back than long; clypeal height about twice diameter of anterior median eyes, corners of clypeus with incised margins that overlie cheliceral boss; chilum weakly sclerotized, divided, composed of two triangular sclerites. Chelicerae vertical, anterior surface with few, erect setae; promargin with three teeth situated at proximal end of fang furrow, most proximal tooth smallest, retromargin with three larger, more distally situated teeth; very short, narrow, Ishaped posterior sclerite present, separating chelicerae at base. Labium short, distally invaginated at middle, reflexed at almost 90 ° angle relative to sternum. Endites rectangular, distally slightly convergent, with anteromedian scopula and anterolateral serrula consisting of single row of teeth. Sternum round­ ed, without extensions to or between coxae, with erect setae; posterior margin not extending between coxae IV.

Leg formula 4123. Typical leg spination pattern (only surfaces bearing spines listed): femora: I d1­1­1, p0­2­1, r1­2­1; II d1­1­1, p1­2­1, r1­2­1; III d1­1­1, p2­1­1, r1­2­1; IV d1­1­1, p2­1­1, r0­2­1; patellae III, IV p0­1­ 0, r0­1­0; tibiae: I, II d1­0­1, p0­1­0, v4­4­2, r0­1­0; III, IV d1­0­1, p0­1­1, v2­2­2, r0­1­ 1; metatarsi: I p1­1­0, v2­2­2, r1­1­1; II p1­ 1­1, v2­2­2, r1­1­0; III p1­2­2, v2­2­1p, r1­ 1­2; IV p1­1­2, v2­2­1p, r1­2­2; tarsi with three claws and claw tufts, superior claws with several weak teeth, most distal teeth largest, inferior claws unarmed; all tarsi with strong ventral scopulae, scopular hairs distinct from those of claw tufts; trochanters notched, posteriors more strongly so than anteriors; males without tibial crack; metatarsi without preening combs. Abdomen without anterior or dorsal scutum; anterior lateral spinnerets large, composed of two articles, distal article with two major ampullate gland spigots and about 15 small piriform gland spigots; posterior median spinnerets composed of one article, those of male small, tubular, those of female triangular, expanded posteriorly, where bearing three large cylindrical gland spigots arranged in triangle (two spigots situated anteriorly, one posteriorly), anterior two cylindrical gland spigots preced­ ed anteriorly by smaller aciniform and/or minor ampullate gland spigots; posterior lateral spinnerets composed of two articles, distal article about one­fifth as long as proximal article, those of female with at least two large cylindrical gland spigots at base, smaller spigots on tip; colulus represented by setae on small lobe.

Male palp with patella not widened, retrolateral tibial apophysis bipartite; subtegulum and tegulum with interlocking processes, median apophysis heavily sclerotized at least distally, embolus bipartite, with deep ushaped division between prolateral prong and copulatory prong, accompanied by hyaline conductor. Female palp with extremely long, dentate claw. Epigynum without anterior hood, relatively flat, with large median septum; spermathecae posterolaterally situated.