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Hypothenemus japonicus ( Niisima, 1910: 10 ) (Cryphalus)

Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You & Jordal, Bjarte H., 2020, Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae), Insect Systematics and Diversity 4 (3), No. 1, pp. 1-81 : 37-38

publication ID

 https://doi.org/10.1093/isd/ixaa002

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scientific name

 Hypothenemus japonicus ( Niisima, 1910: 10 ) (Cryphalus)
status

 

Hypothenemus japonicus ( Niisima, 1910: 10) (Cryphalus) View in CoL .

Hypothenemus javanus ( Eggers, 1908: 215) (Stephanoderes) View in CoL .

= Stephanoderes obesus Hopkins, 1915: 30 View in CoL (syn: Wood, 1957b).

= Stephanoderes brunneus Hopkins, 1915: 31 View in CoL (syn: Bright, 2019.

= Stephanoderes frontalis Hopkins, 1915: 31 View in CoL (syn: Wood, 1954).

= Stephanoderes philippinensis Hopkins, 1915: 31 View in CoL (syn: Wood, 1975b).

= Stephanoderes bananensis Eggers, 1922: 167 View in CoL (syn: Wood, 1972b).

= Stephanoderes kalshoveni Schedl, 1939c: 35 View in CoL (syn: Wood, 1972b).

= Stephanoderes cryphalomorphus Schedl, 1939e: 14 View in CoL (syn: Wood, 1977a).

= Stephanoderes bituberculatus Eggers, 1940a: 126 View in CoL (syn: Wood, 1977a).

= Stephanoderes subagnatus Eggers, 1940b: 101 View in CoL (syn: Wood, 1972b).

= Stephanoderes pistor Schedl, 1951e: 102 View in CoL (syn: Wood, 1976).

= Stephanoderes prosper Schedl, 1951e: 103 View in CoL (syn: Wood, 1976).

Hypothenemus kamathi Beaver, 1995: 16 .

Hypothenemus kraunhiae Murayama, 1950: 50 View in CoL .

Hypothenemus lefevrei ( Schedl, 1952d: 8) (Stephanoderes) View in CoL .

Hypothenemus leprieuri ( Perris, 1866: 194) (Dryocoetes) View in CoL .

= Stephanoderes albipilis Reitter, 1887: 195 View in CoL (syn: Eggers, 1940e).

= Dryocoetes kraussei Wichmann, 1911: 210 (syn: Eggers, 1921). Hypothenemus leptosquamus Bright, 2019: 152 .

Hypothenemus liberiensis ( Hopkins, 1915: 31) (Stephanoderes) View in CoL .

= Stephanoderes theobromae Eggers, 1932: 32 View in CoL (syn: Wood, 1968). Hypothenemus liliputianus Bright, 2019: 152 .

Hypothenemus lineatus ( Eggers, 1927a: 175) (Stephanoderes) View in CoL .

Hypothenemus longipennis ( Eggers, 1935: 305) (Stephanoderes) View in CoL .

Hypothenemus longipilis Schedl, 1952a: 451 View in CoL .

Hypothenemus loranthus ( Schedl, 1942d: 8) (Margadillius) comb. nov. [ Margadillius View in CoL ].

Hypothenemus macrolobii ( Eggers, 1940c: 234) (Stephanoderes) View in CoL .

Hypothenemus madagascariensis Schedl, 1953c: 82 View in CoL .

Hypothenemus magnus ( Eggers, 1924: 102) (Stephanoderes) View in CoL .

Hypothenemus major Browne, 1970: 555 View in CoL .

Hypothenemus malayensis ( Schedl, 1977c: 499) (Lepiceriodes) View in CoL .

Hypothenemus mallyi ( Hopkins, 1915: 32) (Stephanoderes) View in CoL .

= Stephanoderes soussouensis Eggers, 1943b: 74 View in CoL (syn: Wood, 1972b).

Hypothenemus malus ( Schedl, 1957b: 56) (Stephanoderes) View in CoL .

Hypothenemus mangovorus Schedl, 1961b: 134 View in CoL .

Hypothenemus marginatus Johnson nom. nov.

= Periocryphalus sobrinus Wood, 1974a: 22 View in CoL syn. nov.

Remarks: A replacement name is proposed for Periocryphalus sobrinus Wood, 1974 following the generic synonymy of Hypothenemus and Periocryphalus , creating conflict with Stephanoderes sobrinus Schedl, 1965a . The replacement name is an adjective, referencing the prominent posterolateral margin of the elytra. The holotype of Periocryphalus sobrinus Wood, 1974 at BMNH is lost and erroneously replaced with a Cnesinus sp. , which clearly does not match the species description. A paratype at USNM collected from the same location on a different date matches the original and subsequent descriptions of the species. (Labeled ‘ PARATYPE // Periocryphalus // sobrinus // S. L. Wood 1972 //// Periocryphalus //sobrinus//.72 Wood // S. L. Wood // RS/RGS Exp. Brazil // 12′31′S 51′46′W //// R. A. Beaver F03 // 7.XI.1968 //// S. L. Wood Collection //// USNMENT 01448187’.) Since this species is the type of the genus Periocryphalus , the stability of the name-bearing type is critical. Therefore, the single remaining specimen in the type series at USNM is designated as a neotype of Periocryphalus sobrinus

Wood, 1974a, and subsequently, the neotype of Hypothenemus marginatus Johnson.

Hypothenemus marshalli ( Eggers, 1936b: 36) (Stephanoderes) View in CoL .

Hypothenemus mateui ( Schedl, 1965g: 198) View in CoL ( Stephanoderes View in CoL ).

Hypothenemus melanarius ( Schedl, 1953c: 80) (Cosmoderes) View in CoL .

Hypothenemus melasomus ( Lea, 1910: 140) (Cryphalus) View in CoL .

Hypothenemus meridensis Wood, 2007: 506 View in CoL .

Hypothenemus miles ( LeConte, 1878: 433) (Cryphalus) View in CoL .

Hypothenemus minor ( Eggers, 1927a: 178) (Adiaeretus) View in CoL .

Hypothenemus modestus ( Murayama, 1940: 236) (Cryphalus) View in CoL .

Hypothenemus morigerus ( Schedl, 1957b: 57) (Stephanoderes) View in CoL .

Hypothenemus morio ( Eggers, 1940b: 101) (Stephanoderes) View in CoL .

Hypothenemus morosus Schedl, 1965a: 57 View in CoL .

Hypothenemus mozambiquensis Eggers, 1943b: 75 View in CoL .

Hypothenemus mulongensis ( Eggers, 1940c: 235) (Stephanoderes) View in CoL .

Hypothenemus multidentatulus ( Schedl, 1962f: 95) (Stephanoderes) View in CoL . Hypothenemus multidentatus ( Hopkins, 1915: 28) (Stephanoderes) View in CoL . = Stephanoderes ferrugineus Hopkins, 1915: 29 View in CoL (syn: Wood, 1972b). = Stephanoderes nitidifrons Hopkins, 1915: 31 View in CoL (syn: Wood, 1972b). = Hypothenemus hopkinsi Browne, 1963b: 53 View in CoL (syn: Wood, 1972b). Hypothenemus multipunctatus ( Schedl, 1939c: 36) (Stephanoderes) View in CoL . Hypothenemus muticus ( Schedl, 1961b: 136) (Stephanoderes) View in CoL .

Hypothenemus namosianus Browne, 1983a: 78 View in CoL .

Hypothenemus nanellus Wood, 1971: 34 View in CoL .

Hypothenemus nanoparvus Bright, 2019: 154 .

Hypothenemus natalensis ( Schedl, 1941a: 392) (Archeophalus) View in CoL .

Hypothenemus nesiotus Bright, 2019: 154 .

Hypothenemus nigropiceus ( Schedl, 1951c: 20) (Stephanoderes) View in CoL .

Hypothenemus novateutonicus ( Schedl, 1951e: 105) (Ptilopodius) View in CoL comb. nov. [ Ptilopodius View in CoL ].

Hypothenemus obscurifrons Bright, 2019: 155 .

Hypothenemus obscurus ( Fabricius, 1801: 395) (Hylesinus) View in CoL .

= Stephanoderes asperulus Eichhoff, 1872a: 133 View in CoL .

= Stephanoderes cassiae Eichhoff, 1878b: 152 View in CoL .

= Hypothenemus kuennemanni Reitter, 1902: 140 View in CoL (syn: Wood, 1972b).

= Stephanoderes moschatae Schaufuss, 1905: 8 View in CoL (syn: Wood, 1972b). = Stephanoderes rufescens Hopkins, 1915: 29 View in CoL (syn: Wood, 1972b).

= Stephanoderes buscki Hopkins, 1915: 30 View in CoL (syn: Wood, 1972b).

= Stephanoderes amazonicus Eggers, 1934: 78 View in CoL (syn: Wood, 1972b).

= Hypothenemus emarginatus Schedl, 1942d: 11 View in CoL (syn: Wood, 1972b).

Remarks: Bright (2019) noted that Hypothenemus seriatus (Eichhoff, 1872) was a junior synonym of H. obscurus ( Fabricius, 1801) , as it was synonymized by Wood (1954), based on a specimen compared with the type by Eggers, but the synonymy was largely ignored (including by Wood). We consider the synonymy originally proposed by Wood to be rejected. Hypothenemus obscurus is a highly damaging pest which is not present in some areas where H. seriatus is present. We found that the species were sister in the phylogeny. The morphological differences have been well documented ( Vega et al., 2015), and they are genetically distinct ( Mitchell and Maddox 2010, Johnson et al 2018). It is possible that there is much more variation than documented which may make definitive identification by morphology difficult or impossible, but more evidence would be needed to justify grouping the distinct morphotypes as one species, as well as alternative recognition of the pestiferous beetles which devastate the macadamia and other tropical nut industries. The economic and policy implications of the synonymy could hinder the necessary biosecurity measures to prevent the pest species establishing in other areas.

Hypothenemus opacus ( Eichhoff, 1872a: 132) (Stephanoderes) View in CoL .

= Hypothenemus dolosus Wood, 1974a: 21 View in CoL (syn: Bright, 2019). Hypothenemus paradoxus Schedl, 1964c: 45 View in CoL . Hypothenemus parallelus ( Hopkins, 1915: 25) (Stephanoderes) View in CoL . Hypothenemus parasquamosus Bright, 2019: 160 . Hypothenemus parvulosus Bright, 2019: 160 . Hypothenemus parvulus Browne, 1984d: 93 View in CoL . Hypothenemus paulus Bright, 2019: 162 . Hypothenemus perappositus ( Schedl, 1934: 91) (Stephanoderes) View in CoL . Hypothenemus perexiguus Bright, 2019: 162 . Hypothenemus perhispidus ( Eggers, 1927a: 177) (Stephanoderes) View in CoL . Hypothenemus perpunctatus ( Eggers, 1940c: 233) (Stephanoderes) View in CoL . Hypothenemus piaparolinae Johnson, Atkinson and Hulcr, 2016: 418 . Hypothenemus pilosus Hopkins, 1915: 20 View in CoL . Hypothenemus ponticus Bright, 2019: 164 . Hypothenemus praecellens ( Schedl, 1972c: 288) (Stephanoderes) View in CoL . Hypothenemus pubescens Hopkins, 1915: 19 View in CoL .

= Hypothenemus subelongatus Hopkins, 1915: 19 View in CoL (syn: Wood, 1972b).

= Stephanoderes opacifrons Hopkins, 1915: 25 View in CoL (syn: Wood, 1972b).

= Hypothenemus minutissimus Schedl, 1952a: 450 View in CoL (syn: Wood, 1989). Hypothenemus pubipennis ( Eggers, 1935: 305) (Stephanoderes) View in CoL . Hypothenemus puertoricensis ( Bright and Torres, 2006: 409) (Trischidias) . Hypothenemus pullus ( Wood, 1971: 33) (Periocryphalus) comb. nov. [ Periocryphalus View in CoL ]. Hypothenemus pygmaeomorphus Bright, 2019: 166 . Hypothenemus rotundicollis ( Eichhoff,1878a: 385) (Stephanoderes) View in CoL .

= Stephanoderes sculpturatus Eichhoff, 1878a: 385 View in CoL syn. nov. (syn: Eichhoff, 1896).

= Stephanoderes rotundicollis Eichhoff, 1878b: 45 View in CoL syn. nov.

= Stephanoderes sculpturatus Eichhoff, 1878b: 45 View in CoL syn. nov.

= Stephanoderes quercus Hopkins, 1915: 32 View in CoL syn. nov. Hypothenemus rubrithorax Bright, 2019: 168 . Hypothenemus ruficeps Perkins, 1900: 181 View in CoL . Hypothenemus rugifer ( Schedl, 1965d: 9) (Pachynoderes) View in CoL . Hypothenemus ruginosus Wood, 1989: 178 View in CoL .

= Pachynoderes rugifer Schedl, 1977a: 395 View in CoL (syn: Wood, 1989) (Permanently invalid name due to primary homonymy). Hypothenemus rugosipes Wood, 2007: 515 View in CoL . Hypothenemus sambesianus Eggers, 1943b: 74 View in CoL . Hypothenemus sapporoensis ( Niisima, 1910: 3) (Cryphalus) View in CoL . Hypothenemus sassaensis ( Eggers, 1924: 102) (Stephanoderes) View in CoL . Hypothenemus schedli Browne, 1963b: 54 View in CoL .

= Stephanoderes pubescens Schedl, 1942d: 18 View in CoL (syn: Browne, 1963b). Hypothenemus scutiae ( Schedl, 1959c: 708) (Stephanoderes) View in CoL . Hypothenemus seoulensis Choo and Woo, 1989: 58 View in CoL . Hypothenemus seriatus ( Eichhoff, 1872a: 133) (Stephanoderes) View in CoL stat. res.

= Stephanoderes pulverulentus Eichhoff, 1872a: 133 View in CoL (syn: Wood, 1973).

= Stephanoderes vulgaris Schaufuss, 1897: 209 View in CoL (syn: Wood, 1972b).

= Stephanoderes georgiae Hopkins, 1915: 26 View in CoL (syn: Wood, 1972b).

= Stephanoderes minutus Hopkins, 1915: 26 View in CoL (syn: Wood, 1972b).

= Stephanoderes texanus Hopkins, 1915: 26 View in CoL (syn: Wood, 1954).

= Stephanoderes ficus Hopkins, 1915: 27 View in CoL (syn: Wood, 1954).

= Stephanoderes fiebrigi Hopkins, 1915: 27 View in CoL (syn: Wood, 1954).

= Stephanoderes floridensis Hopkins, 1915: 27 View in CoL (syn: Wood, 1954).

= Stephanoderes pini Hopkins, 1915: 27 View in CoL (syn: Wood, 1954).

= Stephanoderes salicis Hopkins, 1915: 27 View in CoL (syn: Wood, 1954).

= Stephanoderes tamarindi Hopkins, 1915: 27 View in CoL (syn: Wood, 1972b).

= Stephanoderes lucasi Hopkins, 1915: 28 View in CoL (syn: Wood, 1954).

= Stephanoderes soltaui Hopkins, 1915: 28 View in CoL (syn: Wood, 1954).

= Stephanoderes virentis Hopkins, 1915: 28 View in CoL (syn: Wood, 1954).

= Stephanoderes nitidipennis Hopkins, 1915: 29 View in CoL (syn: Wood, 1972b).

= Stephanoderes nitidulus Hopkins, 1915: 29 View in CoL (syn: Wood, 1977c).

= Stephanoderes pecanis Hopkins, 1915: 29 View in CoL (syn: Wood, 1954).

= Stephanoderes subopacicollis Hopkins, 1915: 30 View in CoL (syn: Wood, 1977c).

= Stephanoderes niger Hopkins, 1915: 31 View in CoL (syn: Wood, 1972b).

= Hypothenemus robustus Blackman, 1922a: 88 View in CoL (syn: Wood, 1954).

= Hypothenemus cassavaensis Schedl, 1938a: 453 View in CoL (syn: Wood, 1989).

= Stephanoderes hawaiiensis Schedl, 1941b: 112 View in CoL (syn: Wood, 1989).

= Stephanoderes darwinensis Schedl, 1942c: 178 View in CoL (syn: Wood, 1972b).

= Hypothenemus striatulus Schedl, 1942d: 12 View in CoL (syn: Wood, 1989).

= Hypothenemus marovoayi Schedl, 1953c: 81 View in CoL (syn: Wood, 1989).

= Stephanoderes andersoni Wood, 1954: 1045 View in CoL (syn: Wood, 1972b).

= Stephanoderes liquidambarae Wood, 1954: 1046 View in CoL (syn: Wood, 1972b).

= Stephanoderes asperatus Schedl, 1967c: 226 View in CoL (syn: Wood, 1989). Hypothenemus setiferous Bright, 2019: 169 .

Hypothenemus setosus ( Eichhoff, 1868: 391) (Hypoborus) View in CoL .

= Stephanoderes congonus Hagedorn, 1912a: 337 View in CoL (syn: Wood, 1975b).

Hypothenemus simoni ( Reitter, 1887: 194) (Stephanoderes) View in CoL .

Hypothenemus sobrinus ( Schedl, 1965a: 58) (Stephanoderes) View in CoL .

Hypothenemus socialis ( Schedl, 1957b: 57) (Stephanoderes) View in CoL .

Hypothenemus solitarius ( Schedl, 1950a: 24) (Stephanoderes) View in CoL .

Hypothenemus solivagus Bright, 2019: 171 .

Hypothenemus solocis Wood, 1974a: 21 View in CoL .

Hypothenemus sparsedentatus ( Schedl, 1942c: 178) (Stephanoderes) View in CoL . Hypothenemus sparsus Hopkins, 1915: 20 View in CoL .

= Hypothenemus similis Hopkins, 1915: 20 View in CoL (syn: Wood, 1954).

= Stephanoderes tridentatus Hopkins, 1915: 31 View in CoL (syn: Wood, 1954). Hypothenemus spinatus ( Schedl, 1977b: 282) (Miocryphalus) .

Hypothenemus spinicollis ( Schedl, 1965a: 59) (Stephanoderes) View in CoL .

Hypothenemus spinosus ( Schedl, 1979c: 98) (Lepiceroides) View in CoL .

Hypothenemus squamosulus Johnson nom. nov. [ Ptilopodius View in CoL ].

= Ptilopodius squamosus Schedl, 1953b: 294 View in CoL syn. nov. (secondary homonym).

Remarks: A replacement name is proposed for Ptilopodius squamosus Schedl, 1953b after transferring this species from Ptilopodius , since the combination is already occupied by Stephanoderes squamosus Hopkins, 1915 . The new name is the diminutive of the adjective squamosus (= scaly), referring to the scalelike setae on the pronotum and elytra.

Hypothenemus squamosus ( Hopkins, 1915: 26) (Stephanoderes) View in CoL .

Hypothenemus stigmosus ( Schedl, 1951e: 101) (Stephanoderes) View in CoL .

= Stephanoderes garciae Schedl, 1958c: 42 View in CoL (syn: Wood, 1989).

Hypothenemus striatus ( Atkinson, 1993: 422) (Trischidias) .

Hypothenemus styrax ( Schedl, 1942a: 177) (Stephanoderes) View in CoL .

Hypothenemus subacuminatus ( Schedl,1942a: 177) (Stephanoderes) View in CoL . Hypothenemus subterrestris Johnson, Atkinson and Hulcr, 2016: 421 .

Hypothenemus suspectus Wood, 1974a: 22 View in CoL .

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Schedl, K. E. 1953 c. Fauna Madagascariensis - III. 125. Contribution to the morphology and taxonomy of the Scolytoidea. Mem. Inst. Sci. Madagascar, Serie E. 3: 67 - 106.

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Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Curculionidae

Genus

Hypothenemus

Specimens