Cleobulia coccinea (Vell.) L.P. Queiroz, 2020

Queiroz, Luciano Paganucci de & Snak, Cristiane, 2020, Revisiting the taxonomy of Dioclea and related genera (Leguminosae, Papilionoideae), with new generic circumscriptions, PhytoKeys 164, pp. 67-114 : 67

3.1. Cleobulia coccinea (Vell.) L.P. Queiroz comb. nov.

Basionym: Dolichos coccineus Vell., Fl. Flumin.: 321, 1829 [1825]. Ic. 7 pl. 158. 1831. Type: Brazil, Rio de Janeiro, "Habitat silvis, fruticetisque maritimis", Vellozo (lectotype, designated here: plate 158 in Florae Fluminensis vol. 7, Vellozo 1831).Epitype: Brazil, Bahia, Lençóis, L.P. Queiroz et al. 16029 (epitype, designated here: HUEFS! [200008453]; isoepitypes: ALCB! [046364!], RB! [1173635!], US! [3698469]).

Cleobulia multiflora Mart. ex Benth., Comm. Legum. Gen.: 67. 1837. Type: Brazil, Minas Gerais, Martius s.n. (lectotype, designated here: M! [0240673]), syn. nov.

Note.

A link between Dolichos coccineus Vell. and Cleobulia multiflora Mart. ex Benth. was established by Maxwell (1977), who speculated that they could be synonymous. The description provided by Vellozo (1829: 321) is exceedingly brief, but presents some traits characteristic of this species, such as flowers small and perianth purpureum. The illustration provides more elements to confirm its identity as C. multiflora as it shows resupinate flowers with the standard spreading, the wing petals sagittate and much shorter than the others and the pistil with a straight ovary and style upcurved ca. 90°. There are issues regarding the publication dates of several sections of the Florae Fluminensis but the main text in volume 1 (pages 1 to 329) is considered as having been distributed between 7 September to 28 November 1829 and the illustration volumes on 29 October 1831 ( Carauta 1969, 1972; Stafleu and Cowan 1985; Lima 1995), thus predating and having priority over Cleobulia multiflora published by Bentham in 1837. To avoid misinterpretation of the name proposed by Vellozo (1831)), we are designating an epitype with leaf, flowers and fruits.