Maechidius subcostatus Heller, 1895

Telnov, Dmitry, 2020, A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line, European Journal of Taxonomy 721, pp. 1-210 : 115-117

publication ID	https://doi.org/ 10.5852/ejt.2020.721.1127
publication LSID	lsid:zoobank.org:pub:89E62EF8-2E45-4C59-94B7-6A5603E8939B
DOI	https://doi.org/10.5281/zenodo.4344445
persistent identifier	https://treatment.plazi.org/id/1A3787F6-9D6E-922F-FDAE-FA70CCDF2BF2
treatment provided by	Valdenar
scientific name	Maechidius subcostatus Heller, 1895
status

Treatment

Maechidius subcostatus Heller, 1895 View in CoL

Figs 88 View Figs86–89 , 182–183 View Figs 181–192 , 257 View Figs 242–258 , 282 View Figs 278–285 , 348 View Figs 333–348 , 420, 434, 524, 724–725

Type material

Lectotype [herewith designated]

PAPUA NEW GUINEA • ♂; “ K.Wilhelm-Land, Bongu [p, blue label]// Typus [p, pink label]// 10302 [h] // Staatl. Museum für Tierkunde.Dresden [p]”; SNSD.

Paralectotypes

PAPUA NEW GUINEA • 1 ♂; “ Maechidius subcostatus Heller. [h] Determ.K.M.Heller. [p]// K.K. Wilhelm-Land, Bongu [p, bluish label]// Cotypus ♂ [p, pink label]// SYNTYPE Maechidius subcostatus Heller, 1895 labelled by MFNB 2019 View Materials [p, red label]”; ZMHB • 1 ♀; “ K.K. Wilhelm-Land, Bongu [p, bluish label]// Cotypus ♂ [p, pink label] // SYNTYPE Maechidius subcostatus Heller, 1895 labelled by MFNB 2019 View Materials [p, red label]”; ZMHB • 1 ♀; “ K.Wilhelm-Land, Bongu [p, blue label]// 8186 [h]// Staatl. Museum für Tierkunde.Dresden [p]”; SNSD .

New material

PAPUA NEW GUINEA • 2 specs; “ Bongu [h] K. Wilhelmland [p] 8186 // Staatl. Museum für Tierkunde Dresden [p]”; SNSD • 1 spec.; “ K. Wilhelm-Land, Bongu [p]// 8186 [h] // Staatl. Museum für Tierkunde Dresden [p]”; SNSD • 1 ♂; “ PAPUA: Kokoda. 1,300 ft. ix.1933. L.E.Cheesman. B.M.1933-577.”; BMNH • 3 ♂♂; “ PAPUA: Ishurava. 3,000 ft. vii. 1933. L.E.Cheesman. B.M.1933-577 // Maechidius hirtipes det. G.Frey,1967/68.”; BMNH • 1 ♀; “coll. I. R.Sc.N.B. PAPUA NEW GUINEA Madang Province Canopy Mission Baiteta Light XO 25-IV-1996 Leg Olivier Missa ”; IRSN • 1 ♀; “coll. I. R.Sc.N.B. PAPUA NEW GUINEA Madang Province Canopy Mission Baiteta Light AR 41 30-IV- 1996 Leg Olivier Missa ”; IRSN • 1 ♂; “coll. I. R.Sc.N.B. PAPUA NEW GUINEA Madang Province Canopy Mission Baiteta Light AR 41 01-V- 1996 Leg Olivier Missa ”; IRSN • 1 ♀; “coll. I. R.Sc.N.B. PAPUA NEW GUINEA Madang Province Canopy Mission Baiteta Light AR 8 16-VII-1996 Leg Olivier Missa ”; IRSN • 6 specs; “Collection Naturhistorisches Museum Basel// Papua New Guinea Madang Prov. L. Cizek lgt. // Salemben village 145°24’E 4°42’S 16.XII. 2000, 750 m”; NHMB GoogleMaps • 1 spec.; same labels as for preceding; DTC • 2 ♂♂; “PNG, Madang, Baitabag vill., 50m a.s.l., 145°47’E, 5°08’S M. Janda coll., hand collecting 20.iii.2002 // coll. IECA, České Bud ě jovice Czech Republic ”; IECA GoogleMaps .

Remarks

Heller (1895: 1) based his description of M. subcostatus on an unstated number of specimens. The single SNSD specimen labelled “Typus” is herewith designated as the lectotype and the specimen with the identical locality label becomes a paralectotype. Two ZMHB syntypes also become paralectotypes. A lectotype is designated in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other syntypes are discovered in the future. Additional SNSD specimens with slightly different labels are not considered syntypes since they have different ID numbers from the lectotype.

Description

MEASUREMENTS. Total body length 7.70–9.00 (lectotype) to 9.70 (largest paralectotype) mm.

Dorsum uniformly black-brown, venter and appendages brown. Head convex dorsally between compound eyes, opaque dorsally and ventrally. Male labroclypeus ( Fig. 182 View Figs 181–192 ) broadly emarginate anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views. Female labroclypeus similar to as in male ( Fig. 183 View Figs 181–192 ). Anterolateral angles rounded, moderately strongly protruding, raised up at angle of nearly 90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse inconspicuous setae along anterior and lateral margins. Canthus broadly rounded to obtusely angulate in dorsal view. Punctures of frons irregularly hexagonal, large, dense and shallow. Intervening spaces smooth, much smaller than punctures. Head setae very short, appressed to suberect, not or hardly surpassing length of corresponding punctures.Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, opaque dorsally and laterally. Anterior margin of pronotum broadly emarginate to slightly sinuous with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum broadly rounded in dorsal view, deeply emarginate in basal fourth, crenulate in anterior half, denticulate at lateral emargination, smooth to slightly crenulate in emargination area ( Fig. 257 View Figs 242–258 ). Lateral margin of pronotum sinuous in lateral view. Inconspicuous erect seta present between every two crenulae or denticles. Punctures of pronotal disc large, horseshoeshaped, deep and dense. Intervening spaces glossy, distinctly smaller than punctures. Pronotal setae inconspicuous, seta not surpassing length of median ‘bulb’ of horseshoe-like puncture. Hypomeron separated from prosternum by high nearly straight carina, its anterolateral margin slightly angulate medially, with long setae on anterolateral margin opposite to compound eye ( Fig. 282 View Figs 278–285 ). Antennal pocket moderately deep. Scutellar shield narrowly rounded apically. Elytron opaque, with vague tracks of four variously broadly interrupted glabrous longitudinal carinae, including sutural one. Elytral punctures linear (elongate and narrow), incision-shaped, moderately deep. Intervening spaces densely microreticulate ( Fig. 348 View Figs 333–348 ). Appressed to suberect seta rises from anterior margin of each incision, not surpassing length of corresponding incision. Disc and lateral margins of elytra in part covered with microscopical velvety pubescence. Male abdomen with sparse clavate scale-like golden setae ( Fig. 434 View Figs 424–435 ). Male pygidium slightly convex dorsally, with large shallow dense annular punctures, female pygidium flattened dorsally. Intervening spaces microreticulate, distinctly smaller than punctures. Male and female pygidium flattened dorsally, opaque, with dense large and shallow annular punctures (Fig. 524). Setae of pygidium inconspicuous, short and suberect, in anterior part not, in distal part surpassing length of corresponding punctures. Male protibia slightly widened distally, with three external teeth: distal one largest and narrow, median broad and obtuse and basal much smaller, strongly obtuse (Fig. 420). Male protibial terminal spur short and straight, female one longer and stronger pointed. Male metatibial terminal spurs long, unequal, lower spur slightly curved, female one more strongly unequal and straight. Tarsal claws with pulvilli. Underside of male basal metatarsomere with a brush of long dense setae. Aedeagus as in Figs 724–725 View Figs 712–725 .

Sexual dimorphism

Female generally more robust, with flattened pygidium, comparatively shorter antennal lamellae and longer protibial terminal spur and more strongly unequal metatibial terminal spurs.

SNSD	SNSD
MFNB	Italy, Udine, Museo Friulano di Storia Naturale
ZMHB	Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet
BMNH	United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]
IRSN	Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique
DTC	Dmitrijs Telnovs
R	Departamento de Geologia, Universidad de Chile
IRSN	Institut Royal des Sciences Naturelles de Belgique
NHMB	Natural History Museum Bucharest
IECA	Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology