Platyderus (Eremoderus) weiratheri Maran , 1940

Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice, 2022, Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini), Deutsche Entomologische Zeitschrift 69 (1), pp. 71-122 : 71

publication ID	https://dx.doi.org/10.3897/dez.69.83840
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scientific name	Platyderus (Eremoderus) weiratheri Maran , 1940
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Treatment

2. Platyderus (Eremoderus) weiratheri Maran, 1940

Figs 1B View Figure 1 , 5A View Figure 5 , 7B View Figure 7 , 8B View Figure 8 , 10B View Figure 10 , 12B View Figure 12 , 13A View Figure 13 , 16 View Figure 16

Platyderus weiratheri Mařan, 1940: 25 (type locality: "Lydien in montibus Tmolos" [= Bozdağlar], Turkey).

Note on type locality.

The mount Bozdağlar (Ancient Greek name: Tmōlos), with maximal elevation 2159 m a.s.l., is situated in the southwestern part of the Anatolian Peninsula.

References.

Platyderus weiratheri : Jedlička 1963: 22; Lorenz 1998: 375; Hovorka and Sciaky 2003: 523; Lorenz 2005: 396; Hovorka 2017: 760.

Type material.

Should consist of two male and three female specimens ( Mařan ibid.) all of them syntypes according to Art. 73.2.1. (ICZN 1999). From these specimens, we found only one male syntype with extracted genitalia, here designated as lectotype. This specimen and its genitalia (with missing urite) are glued on two separate white cards on the same pin. The specimen designated herewith as lectotype is labelled as follows: '♂ [w, p] // Tmolos-Gbg., Lydien, / West-Klainasien. / Weirather, Innsbruck [w, p] // TYPUS [r, p] // Mus. Nat. Pragae / Inv. 927 [o, h/p] // Platyderus weiratheri m. Dr. Mařan det. [w, h&p] // lectotype label’ [NMPC]. The depositories and present state of remaining four type specimens remain unknown.

Other material examined.

Turkey: Bozdağlar (= Mount Tmolus) situated on territories of districts of İzmir, Manisa and Aydin: 1♂, ' Tmolos-Gbg., Lydien, / West-Kleinasien. / Weirather, Innsbruck // Platyderus spec.? det. J. Müller (vergl. mit cyprius!) // Coll. Mus . / Vindob.' (NMW); 1♂, 1♀, ' Tmolos-Gbg., Lydien, / West-Kleinasien. / Weirather, Innsbruck // Coll. Mus . / Vindob.' (NMW); 2♂♂, 1♀, ' Tmolos-Gbg., Lydien, / West-Kleinasien. / Weirather, Innsbruck’ (NMW) ; 1♂, ' Boz dağ, vt N / 1150-1200 m / 5-5-1995 // TURQUIE / Izmir / C. JEANNE // Collection / Machard’ (cMAC); 3♀♀, ʻTURKEY, vil. Izmir , Boz / dağlari, Boz dağ köy env. / 1500-1700 m a.s.l. / 30.5.-3.6.̍03, R.Lohaj lgt.' (NMNHS, cWR).

TME: 11 specimens. TGE: 2♂♂, 1♀.

Diagnosis.

See “Diagnosis” of P. chatzakiae sp. nov.

Redescription.

Habitus. Specimens of large size for Platyderus species (BL: 7.00-8.10 mm; BW: 2.35-2.60 mm), with rather elongate, moderately convex body (Fig. 1B View Figure 1 ). Measurements and ratios. See Table 1 View Table 1 . Color and lustre. Body dorsally and ventrally dark reddish, head and pronotum mostly lighter than elytra, appendages (antennae, palpi and legs) rufous, lighter colored than dorsal surface. Integument moderately shiny, head and pronotum slightly shinier than elytra. Microsculpture and punctation. Pronotum without regular microreticulation. Elytral intervals and scutellum with distinct isodiametric sculpticells; basal margin and lateral gutter with such sculpticells less evident or without microsculpture. Ventral surface with hardly visible isodiametric (proepisternum) or slightly transverse sculpticells (abdominal ventrites at sides) or mostly sculpticells not apparent. Head with some punctures and wrinkles only clypeus and frons, labrum, disc and vertex without punctures and wrinkles. Pronotum in basolateral part coarsely and sensely punctate, basal punctures reaching or not reaching the anterior half laterally, disc and anteroapical part mostly smooth, anterior third medially finely punctate and wrinkled. Elytral intervals mostly with a row of punctures, rarely without punctation. Abdominal ventrites 1-5 wrinkled at sides. Head. One-third or narrower than pronotum wide (PW/HW= 1.32-1.44). Eyes slightly convex. Labrum subrectangular, slightly shorter than clypeus, with anterior margin concave. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows barely distinct, small and shallow. Paraorbital sulci straight, moderately deep, ending backwards slightly before level of posterior supraorbital pore. Thorax. Pronotum about one eighth to one quarter wider than long (PW/PL= 1.13-1.27), with widest point at second quarter. Anterior and posterior transverse impressions slightly distinct medially. Sides sinuate, convex medially and anteriorly, slightly to hardly concave posteriorly; anterior border lack in medial 1/10. Metepisternum about as long as wide, MA/MI= 0.95-1.02. Elytra. Elongate, about two thirds longer than wide (EL/EW= 1.63-1.71), two and a half times or longer than pronotum (EL/PL= 2.55-2.83), and a third wider than pronotum (EW/PW= 1.29-1.40), with widest point at medial third or third quarter. Parascutellar striole and striae well-impressed, more coarsely punctate than in P. chatzakiae sp. nov. and less coarsely than in P. vanensis sp. nov.; striole short, not joining stria 1; striae 1-6 reaching or almost reaching basal bead, 7 reduced before. Interval 3 with three discal setiferous punctures on each elytron. Umbilicate setiferous series consisting of 15-16 (rarely 17) punctures on each elytron. Legs. Posterior side of profemur with one seta in basal third and one seta in medial third. Anterior side of mesofemur ventrally mostly with 4 equidistant setiferous punctures, rarely with 3 or 5 punctures (Fig. 5A View Figure 5 ). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third. Male genitalia. Urite IX with proximal margin almost asymmetrical (Fig. 7B View Figure 7 ). Median lobe of aedeagus in lateral view as in P. chatzakiae , with shaft little longer (Fig. 8B View Figure 8 ); median lobe in ventral view straight, about 3.5 times longer than wide (Fig. 10B View Figure 10 ); apical lamella (dorsal view) as in P. chatzakiae . Internal sac as in P. chatzakiae , only ventral sclerite narrowed toward both proximal and distal ends, with widest point in middle (Fig. 10B View Figure 10 ). Right paramere on Fig. 12B View Figure 12 . Female genitalia (Fig. 13A View Figure 13 ). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.

Habitat.

Nothing is known about bionomics of this species.

Distribution.

Apparently endemic to the mountains of the Bozdağlar Mountain, in Southwest Turkey (Fig. 16 View Figure 16 ).

" Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis " group

Diagnosis. Includes species with basal foveae of pronotum and adjacent lateral areas coarsely and densely punctate with punctures at sides usually reaching anterior half. The prosternum laterally and proepisternum also coarsely and densely punctate. The ventral sclerite of internal sac of median lobe in ventral view is elongate, relatively wide, with straight distal end; the same in lateral view is significantly widened anteriorly having maximum width at the distal third about three or more times larger than width at the proximal third (Figs 8C-F View Figure 8 , 10C-F View Figure 10 ). Most specimens lack also the first elytral discal puncture and have apical gonocoxite with proximal dorsolateral ensiform seta finer than the distal seta.

In addition, the Iranian congeners of the " Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis " group differ from those of the " Platyderus davatchii " group in: (1) sides of pronotum nearly straight to the posterior angles (vs. sides of pronotum concave to the posterior angles) and (2) pronotum appreciably wider than long (PW/PL>1.20, vs. PW/PL <1.20). For further differences between taxa of the group dealt with here and P. lassallei see “Diagnosis” under the " Platyderus Eremoderus lassallei " species group.

Habitat. The habitats of the Eremoderus -species in the Zagros (Iran) consist mostly of subalpine slopes, partly near snowfields, and seldom of cultivated grasslands and small orchards. Outside the cultivated grasslands along the rivers and streams the vegetation is xerophilic. The altitude of the area where P. felixi sp. nov. was collected is appreciably lower than that of the localities of P. iranicus sp. nov.

" Platyderus Eremoderus iranicus " subgroup

Notes. This complex includes two allopatric, high altitudinal species, P. felixi and P. iranicus with parascutellar striole, striae 1-8 scarcely punctate and moderately impressed and quite short and crooked right parameres (Fig. 12C, D View Figure 12 ).