Tylopus nodulipes ( Attems, 1953 )

Nguyen, Anh D., 2012, Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species, Raffles Bulletin of Zoology 60 (2), pp. 289-311 : 301-303

Tylopus nodulipes ( Attems, 1953)

( Figs. 13 , 18 )

Agnesia nodulipes Attems, 1953: 174 , Figs. 70–74; Jeekel, 1965: 98

Tylopus nodulipes :— Jeekel, 1968: 60; Golovatch, 1984: 70, Fig. 27; Golovatch & Enghoff, 1993: 106, Figs. 71–74; Enghoff, Golovatch & Nguyen, 2004: 40

Material examined. — 1 male, 2 females (IEBR-91), Lao Cai Province, Van Ban District, Nam Xay commune (22°05'N, 104°05'E), primary forest, 1000 m a.s.l., coll. Nguyen Duc Anh, 31 Mar.2005 – 15 Apr.2005 ; 1 male (IEBR-104), Ha Tinh Province, Huong Son District , Son Tay commune, secondary forest, 600 m a.s.l., coll. Nguyen Duc Anh, 3 May 2004 ; 4 males, 5 females (IEBR-105), Ha Tinh Province, Huong Son District , Son Tay commune, secondary forest, 600 m a.s.l., coll. Nguyen Duc Anh, 25 May 2004 ; 2 males, 2 females (IEBR-153), Nghe An Province, Pu Mat National Park (18°46'– 19°12'N, 104°24'– 104°56'E), Chem Waterfall, secondary forest, 430 m a.s.l, coll. Nguyen Duc Anh, 4–10 Apr.2011 ; 1 male, 1 female (IEBR-154), same date as sample IEBR-153.

Remarks. — The specimens examined agree well with the original description, but their body is larger (metazonal width 3.3-3.7 mm vs. 3.0 mm in the original description). T. nodulipes has been known from Luong Prabang ( Laos) and Fanxifan Mt. ( Vietnam) ( Attems, 1953), and the new records presented here suggest that this species is distributed widely in the northern part of Vietnam.

Figures

Fig. 13. Tylopus nodulipes (Attems, 1953) from Van Ban Distr., Lao Cai: right gonopod, mesal view (A) and lateral view (B); postfemoral region, mesal view (C) and lateral view (D).

Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach Ma–Ba Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi Doup–Nui Ba NP (T. spinisterna).