Dacrysoma ravelosoni, Darby, Michael, 2013

Darby, Michael, 2013, Studies of Madagascan Ptiliidae (Coleoptera) 1: The Tribe Discheramocephalini including eighteen new species, Zootaxa 3718 (3), pp. 239-266 : 256-257

Dacrysoma ravelosoni sp. n.

(Figs. 10, 39, 44, 56, 65, 82, 83, 94)

Habitus Fig. 10. Length 0.94–0.96 mm. Colour dark brown; legs and antennae dusky yellow. Antennae very long, antennomeres 3–9 very narrow and parallel sided, Fig. 44 . Pronotum Figs. 56 , 65 ; 0.24 mm long, 0.32 mm wide, lateral margins slightly angled. Elytra 0.63 mm long, 0.50 mm wide, foveae finer and denser than pronotum, tending to form longitudinal rows.

Male: aedeagus Fig. 94.

Female: micropterous/apterous. Spermatheca shows some variation in length, Figs. 82, 83

Etymology. Named in honour of R. Raveloson who collected many of the specimens.

Remarks. The large size, long thin antennae, dark colour and presence of micropterous/apterous females, will quickly distinguish this species.

Differences in length of the spermatheca and tip of the aedeagus suggested that two species might be involved, but the presence of intermediate forms ultimately discouraged this view.

Type data. Holotype: ♂, Ambohitantely Spec. Res., ABT/08/2011, Forest litter forest edge, sifting, Winkler app. extr., S.18˚11’51.8’’ E. 47˚17’03.5’’, 1523 m, 19.iv.2011, P.Banar & R. Raveloson (BMNH).

Paratypes: all records by Winkler extraction of sieved samples collected in the Ambohitantely Spec. Res. unless stated otherwise. 17 ♀ 9 ♂, same data as holotype; 1 ♀ 3 ♂, ABT/10/2011, sifting plant residues under Pandanus , S.18˚11’39.2’’ E.47˚17’12.6’’. 1674 m, 19.iv.2011, L.S.Rahanitriniaina & R.Raveloson (BMNH); 10 ♀ 2 ♂, ABT/ Nov/2011/08, forest litter, S.18˚11’48.5’’ E. 47˚17’09.3’’, 1595 m, 18.xi.2011, P. Banar (BMNH); 1 ♂ 1 ♀, ABT/ Nov/2011/11, forest litter, S.18˚10’52.6’’ E.47˚17’22.5’’, 1497 m, 18.ix.2011, L.S.Rahanitriniaina (BMNH); 5 ♀, 1 ♂, ABT/ Sept/2011/01, forest litter under Pandanus , S.18˚11’45.1’’ E.47˚17’13.4’’, 1604 m, 1.ix.2011, L.S.Rahanitriniaina (BMNH); 6 ♀ 2 ♂, ABT/ Nov 2011/18, litter under group of palm trees, S.18˚11’48.7’’ E.47˚17’11.3’’, 1614 m, 22.xi.2011, L.S.Rahanitriniaina (BMNH); 4 ♀, ABT/19/2011, litter and plant residues under Pandanus , S.18˚11’44.8’’ E.47˚17’14.2’’, 1609 m, 20.iv.2011, L.S.Rahanitriniaina & R.Raveloson (BMNH); 3 ♂ 1♀, ABT/ Nov/2011/13, forest litter, S.18˚10’56.9’’ E.47˚17’21.9’’, 1497 m, 18.xi.2011, L.S.Rahanitriniaina (BMNH); 12 ♀ 1 ♂, ABT/12/2011, plant residues under palm tree, S.18˚11’48.9’’ E.47˚17’10.5’’, 1602 m, 19.iv.2011, L.S.Rahanitriniaina & R.Raveloson (BMNH); 1 ♂, ABT/ Nov/2011/16, forest litter under Pandanus , S.18˚11’42.6’’ E.47˚17’10.8’’, 1620 m, 21.xi.2010, L.S.Rahanitriniaina (BMNH); 1 ♂, ABT/03/2011, wet aluvia by stream, S.18˚10’51.7’’ E.47˚17’21.6’’, 1593 m, 19.iv.2011, L.S.Rahanitriniaina (BMNH); 3 exs, ABT/ Nov/2011/10, forest litter under palm tree, 1600 m, 18.xi.2011, P. Banar (BMNH); 2 ♂ 2 ♀, ABT/10/2011, plant residues under Pandanus , S.18˚11’39.2’’ E.47˚17’12.6’’, 1674 m, 19.iv.2011, L.S.Rahanitriniaina & R.Raveloson (BMNH); 32 exs, ABT/ Nov/2011/01, forest litter under big Pandanus , S.18˚11’52.7’’ E.47˚17’0.4’’, 1603 m, 16.xi.2011, L.S.Rahanitriniaina & P. Banar (BMNH); 5 ♀ 4 ♂, ABT/18/2011, forest litter, S.18˚11’43.9’’ E.47˚17’6.1’’, 1613 m, 16.xi.2011, L.S.Rahanitriniaina & R.Raveloson (BMNH); 2 ♂ 1 ♀, ABT/17/2011, forest litter, S.18˚11’43.1’’ E.47˚17’19.3’’, 1636 m, 20.iv.2011, L.S.Rahanitriniaina & R.Raveloson (BMNH); 1♀, ABT/ Nov.2011/09, forest litter, 1610 m, 22.xi.2011, L.S.Rahanitriniaina (BMNH); 1♂ 1♀, ABT/ Nov.2011/18, litter under group of plam trees, S.18˚11’48.7’’ E.47˚17’11.3’’, 1614 m, 22.xi.2011, L.S.Rahanitriniaina (BMNH); 1 ex, ABT/ Sept.2011/17, forest litter, S.18˚11’04.8’’ E.47˚17’11.8’’, 1598 m, 4.ix.2011, L.S.Rahanitriniaina (BMNH); 7♀♀, ABT/ Sept.2011/15, forest litter, S.18˚10’57.3’’ E.47˚17’17.2’’, 1565 m, 3.xi.2011, L.S.Rahanitriniaina (BMNH); 30 ♀ 6 ♂, ABT/ Nov.2011/21, forest litter under Pandanus , S.18˚11’48.5’’ E.47˚17’06.1’’, 1656 m, 25.xi.2011, L.S.Rahanitriniaina (BMNH); 1♀, ABT/ Sept.2011/29, forest litter under Pandanus , S.18˚11’54.2’’ E.47˚17’05.5’’, 1613 m, 6.ix.2011, L.S.Rahanitriniaina (BMNH); 1 ♀, ABT/ Sept.2011/03, forest litter under Pandanus , S.18˚11’46.4’’ E.47˚17’09.9’’, 1570 m, 1.ix.2011, L.S.Rahanitriniaina (BMNH); 4 ♀ 2 ♂, forest litter (bamboo, Pandanus ), S.18˚10’52.3’’. E.47˚17’21.3’’, 1493 m, 2.ix.2011, L.S.Rahanitriniaina (BMNH); 2 ♀, ABT/ Nov.2011/03, forest litter under Pandanus , S.18˚11’02.2’’ E.47˚17’13.4’’, 1565 m, 4.ix.2011, L.S.Rahanitriniaina (BMNH); 1 ♀, ABT/ Sept.2011 / 0 8, forest litter, S.18˚11’42.9’’ E 47˚17’20.3’’, 1640 m, 17.xi.2011, L.S.Rahanitriniaina (BMNH); 1 ♂, ABT/ Nov/ 2011/03, forest litter, S.18˚11’42.9’’ E 47˚17’20.3’’, 1640, 17.xi.2011, 1640, 17.xi.2011, L.S.Rahanitriniaina (BMNH); 5 ♂ 3 ♀, ABT/ Sept.2011/11, forest litter under palm tree close by waterfall, S.18˚.10’.58.3’’ E.47˚.17’.22.2’’, 1518 m, 3.ix.2011, L.S.Rahanitriniaina (BMNH); 7 ♀ 8 ♂, ABT/ Sept.2011/13, forest litter under Pandanus and by big tree, S.18˚10’58.4’’ E.47˚17’22.6’’, 1525 m, 3.ix.2011, L.S.Rahanitriniaina (BMNH); 4 ♀, ABT/ Sept.2011/27, forest litter under palm trees, S.18˚11’53.3’’. E.47˚17’11.2’’, 1602 m, 6.ix.2011, L.S.Rahanitriniaina (BMNH); 10 exs, ABT/ Nov.2011/17, forest litter wet material close to stream, S.18˚11’48.6’’ E.47˚17’05.7’’,[no altitude], 21.xi.2011, P. Banar (BMNH); 6 ♀, ABT/ Sept.2011/04, forest litter under Pandanus , S.18˚11’46.4’’ E.47˚17’09.9’’, 1570 m, 1.ix.2011, L.S.Rahanitriniaina (BMNH); 33 exs; ABT/ Sept.2011/04, forest litter by stream, S.18˚10’52.3’’ E.47˚17’21.3’’, 1500 m, 2.ix.2011, L.S.Rahanitriniaina (BMNH); 6 ♀, ABT/ Nov.2011/12, forest litter, S.18˚10’53.3’’ E.47˚17’22.4’’, 1482 m, 18.xi.2011, L.S.Rahanitriniaina (BMNH); 2 ♀ 1 ♂, ABT/ Sept.2011 /, forest litter under big rock by stream, S.18˚10’52.5’’ E.47˚17’21.2’’, 1476 m, 2.ix.2011, L.S.Rahanitriniaina (BMNH); 2 ♂, ABT/ Sept.2011/09, forest litter under Pandanus close to river, S.18˚10’53.5’’ E.47˚17’22.3’’, 1490 m, 2.ix. 2011, L.S.Rahanitriniaina (BMNH); 2 ♂, ABT/01/2011, forest litter under Pandanus very dry material, S. 18 10 51.7 E. 47 17 21.6, 1593 m, 19.iv.2011, P. Banar & R. Raveloson (BMNH); 3 ♂ 3 ♀, ABT/ Nov/2011/19, forest litter, S.18˚10’56.9’’ E. 47˚21’17.9’’, 1497 m, 23.xi.2011, L.S.Rahanitriniaina (BMNH); 4 exs, ABT/05/2011, forest litter by waterfall, S. 18o10’53.6’’ E.47o17’22.4’’, 1503 m, 19.iv.2011, L.S.Rahanitriniaina (BMNH); 2 exs, ABT/ Sept.2011/08, forest litter, S.18˚11’42.9’’ E.47˚17’20.3’’, 1640 m, 17.xi.2011, L.S.Rahanitriniaina (BMNH); 4 exs, ABT/01/2011, forest litter under Pandanus very dry material, S.18˚10’51.7’’ E.47˚17’21.6’’, 1593 m, 19.iv.2011, P. Banar & R. Raveloson (BMNH);); 5 exs, ABT/ Nov.2011/12, forest litter, S.18˚10’53.3’’ E.47˚17’22.4’’, 1482 m, 18.xi.2011, L.S.Rahanitriniaina (BMNH); 3 exs, Ankarafantsika N.P.,AKF/ 05/2011, forest litter, S.16˚ 18’ 05.1’’ E. 46˚ 49’ 08.3’’, 89 m, 23.iv.2011, L.S.Rahanitriniaina & R. Raveloson (BMNH).

Figures

FIGURES 41 – 55. Dacrysoma spp. antennomeres 3 – 11 (hairs omitted). 41, D. ranomafanense; 42, D. varium; 43, D. denticulatum; 44, D. ravelosoni; 45, D. rahanitriniainae; 46, D. dolorosum; 47, D. nigerrimum; 48, D. fabrum; 49, D. imitatum; 50, D. longulum; 51, D. subinsulsum; 52, D. andasibense; 53, D. fusum; 54, D. pilosum; 55, D. delicatum.

FIGURES 56 – 61. Dacrysoma spp. pronota. 56, D. ravelosoni (x 515); 57, D. nigerrimum (x 530); 58, D. ranomafanense (x 555); 59, D. fabrum (x 555); 60, D. varium (x 525); 61, D. rahanitriniainae (x 635).

FIGURES 62 – 76. Dacrysoma spp. pronota. 62, D. ranomafanense; 63, D. varium; 64, D. denticulatum; 65, D. ravelosoni; 66, D. rahanitriniainae; 67, D. dolorosum; 68, D. nigerrimum; 69, D. fabrum; 70, D. imitatum; 71, D. longulum; 72, D. subinsulsum; 73, D. andasibense; 74, D. fusum; 75, D. pilosum; 76, D. delicatum.

FIGURES 77 – 89. Dacrysoma spp. spermathecae. 77, D. ranomafanense; 78, D. varium; 79, D. delicatum; 80, D. denticulatum; 81, D. rahanitriniainae; 82, 83, D. ravelosoni; 84, D. pilosum; 85, D. nigerrimum; 86, D. fabrum; 87, D. andasibense; 88, D. imitatum; 89, D. subinsulsum.