Pupina perakensis Moellendorff , 1891

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina perakensis Moellendorff, 1891

Figs 29G , 34D

Pupina arula var. perakensis Möllendorff, 1891: 345. Type locality: Bukit Pondong, Perak [Gunung Pondok, Perak State, Malaysia].

Pupina arula perakensis - van Benthem Jutting 1949: 58, Cameron Highlands, Pahang; Telom Valley, near Gunong Siku, Pahang; Kuala Legap, Plus Valley, Perak [Malaysia]. van Benthem Jutting 1960: 13, hill near the hot springs, ca. 400 m from the main road from Tandjong Rambutan to Ipoh, near Tambun, Perak. Maassen 2001: 40.

Pupina (Tylotoechus) arula perakensis - Laidlaw 1928: 34. Zilch 1957: 44, pl. 2, fig. 17.

Pupina lowi [non Morgan]- Foon et al. 2017: 40, 41, fig. 15d, Ipoh, Perak.

Pupina tchehelensis [non Morgan]- Foon et al. 2017: 41, fig. 16a, Ipoh, Perak.

Type material examined.

Lectotype SMF 109969/1 (Figs 29G , 34D ) from Bukit Pondong, Perak.

Diagnosis.

Shell fusiform; last whorl ca. 70% of shell height. Apertural lip thickened but not expanded; apertural lip curved when observed from lateral view. Parietal callus and parietal tooth highly thickened; columellar tooth curved downward like an earlobe.

Differential diagnosis.

Pupina perakensis is most similar to P. crosseana , but differs in parietal callus and parietal tooth very thickened, and a columellar tooth curved downward like an earlobe.

Distribution.

Perak and Pahang States, Malaysia ( Maassen 2001).

Remarks.

This taxon has always been treated as a subspecies of P. arula ( van Benthem Jutting 1949; Zilch 1957; Maassen 2001). However, it is different from P. arula in having a glossy shell surface, a more fusiform shape with a higher spire; and a less bulging last whorl; additionally, the occurrence of this taxon is ca. 1,800 km from that of P. arula . Thus, this taxon is herein elevated to the specific level.

By comparing with the type specimen, the specimen of P. tchehelensis figured in Foon et al. (2017: fig. 16a) from Gunung Tempurung Plot 2, Ipoh, Perak should belong to P. perakensis (Foon, pers. comm.). Although the P. lowi specimen figured in Foon et al. (2017: 15d) from Bat Cave Hill, Ipoh, Perak has a shorter spire, we preliminarily identify this specimen as P. perakensis as well due to an overall character in the Pupina arula species group, a similar glossy surface to the type specimen, and its nearby locality to the type locality.

Figures

Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J).

Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 " forma minor " from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B).