Ptomaphagus (s. str.) nepalensis Perreau, 1988

Wang, Cheng-Bin, Perreau, Michel, Ruzicka, Jan & Nishikawa, Masaaki, 2017, Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae), ZooKeys 715, pp. 69-92 : 73-75

Ptomaphagus (s. str.) nepalensis Perreau, 1988 Figs 1 A–B; 2 A–J; 3 A–F

Ptomaphagus (s. str.) nepalensis Perreau, 1988: 1005 ( Ptomaphagus ; type locality: Népal, district de Lalitpur, Phulcoki [ca. 27°34'N 085°25'E], 2600 m; MHNG); Perreau, 2000: 363 ( Ptomaphagus (s. str.); in catalog); Perreau, 2004: 178 ( Ptomaphagus ( Ptomaphagus ); in catalog); Perreau, 2015: 249 ( Ptomaphagus (Ptomaphagus) ; in catalog).

Material examined.

Type material. Paratypes: 1♂, NEPAL, Kath- / mandu District // Phulcoki 2600 m / 20.IV.1982 / A. & Z. Smetana // PARATYPE (MHNG); 1♂, NEPAL Lalitpur / Distr. Phulcoki / 2600 m 16.X.[19]83 / Smetana & Löbl // Ptomaphagus / nepalensis / Perreau 1988 // PARATYPE // MHNG / ENTO / 00003344 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2600 m 14.X.[19]83 / Smetana & Löbl // PARATYPE // MHNG / ENTO / 00003345 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2650 m 15.X.[19]83 / Smetana & Löbl // PARATYPE // MHNG / ENTO / 00003346 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2650 m 14.X.[19]83 / Smetana & Löbl // PTOMAPHAGUS / NEPALENSIS n. sp. / M. PERREAU det. 1987 // PARATYPE // MHNG / ENTO / 00003347 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2700 m 15.X.[19]83 / Smetana & Löbl // PARATYPE (MHNG); 1♂, NEPAL, Kath- / mandu District // Phulcoki 2600 m / 20.IV.1982 / A. & Z. Smetana // PARATYPE // MHNG / ENTO / 00003343 (MHNG); 1♀, NEPAL, Kath- / mandu District // Phulcoki 2600 m / 22.IV.1982 / A. & Z. Smetana // PARATYPE // MHNG / ENTO / 00003342 (MHNG); 2♂♂, NEPAL: distr. / Kathmandu: Phulcoki / 2500 m, 28-29.IV.[19]84 / Löbl - Smetana (CMPR).

Redescription.

Male. EBL: 2.9-3.0 mm. Length of different body parts: HL: AL: PL: ELL = 0.5: 0.9: 0.7: 1.6 mm; width: HW: EW: PW: ELW = 0.8: 0.1: 1.1: 1.1 mm. Proportion of antennomeres from base to tip in μm (length × width): 132 × 54, 89 × 50, 69 × 50, 45 × 52, 47 × 60, 38 × 72, 74 × 89, 29 × 90, 67 × 97, 75 × 103, 137 × 94.

Habitus (Fig. 1A) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, basal three or four antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi more or less yellowish. Dorsum continually clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.

Head transverse, HW/HL = 1.5. Clypeofrontal suture absent. Clypeus with anterior margin almost straight. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 2A) slender, AL/HW = 1.1; antennomere III shorter than II; VI with length/width = 0.5; XI pear-shape.

Pronotum (Fig. 2B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, gradually narrowing from posterior to anterior; hind angles slightly pro jected backwards and subacute. Posterior margin widely but shortly protruded in middle part, emarginate near hind angles.

Elytra oval, widest at about basal 2/7, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 2G) rounded. Sutural striae present. Metathoracic wings fully developed.

Prolegs robust, with basal three protarsomeres (Fig. 2C) expanded: TW/BTW = 1.3. Protibiae (Fig. 2E) expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender, straight, but relatively short.

Abdominal ventrite VIII (Fig. 2I) distinctly emarginate at posterior edge, and much deeper at median. Spiculum gastrale of genital segment (Fig. 2J) with about 2/5 of length protruding beyond anterior edge of epipleurite IX.

Aedeagus (Fig. 3A) rather long and slender, with median lobe gradually narrowing towards a lanceolate apex and terminated to a widely subrounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 3C) inserted with two ventrally oriented setae on the left side and three ventrally oriented setae on the right side; parameres narrow, reaching about apical 1/7 of median lobe, each apex (Fig. 3D) with two long lateral setae and one shorter apical seta. In lateral view (Fig. 3B), median lobe slender, regularly bent ventrally, and gradually tapering to a round apex. Endophallus with stylus quite slender, a transverse nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.

Female. Similar to male in general appearance (Fig. 1B), including elytral apices (Fig. 2H), but distinguished by the following characteristics: protarsi (Fig. 2D) simply linear; protibiae (Fig. 2F) narrower; abdominal ventrite VIII (Fig. 3E) almost rounded at posterior edge and slightly protruded at median; genital segment and ovipositor as shown in Fig. 3F; spermatheca (Fig. 3F) sinuous or coiled in distal part, not coiled in proximal part.

Distribution.

Nepal.