Glossanodon cf. leioglossus (Valenciennes, 1848)

Brzobohatý, Rostislav & Nolf, Dirk, 2018, Revision of the Middle Badenian fish otoliths from the Carpathian Foredeep in Moravia (Middle Miocene, Czech Republic), Cybium 42 (2), pp. 143-167 : 148

Glossanodon cf. leioglossus (Valenciennes, 1848) ( Pl. 1 ,

Fig. 5a, b). A perfectly preserved otolith from Lomnička (length = 2.5 mm; height = 1.6 mm) could be conspecific with the present-day species G. leioglossus , but more material is required for an unambiguous specific identification. This Eastern Atlantic fish (including the Western Mediterranean and southern part of the Adriatic Sea) can be found near the bottom on the outer shelf and upper slope ( Jardas, 1996). G. cf. leioglossus represents the first otolith-based fossil record of the genus, but skeletons of the fossil G. musceli (Paucá, 1929) are relatively abundant in the Lower Oligocene of the Carpathians (e.g. Kotlarczyk et al., 2006).

Nansenia sp. ( Pl. 1 , Fig. 12a, b). Otoliths of the genus Nansenia are rare in the European Neogene and are usually mentioned as Nansenia sp. ( Nolf and Steurbaut, 1983; Schwarzhans, 1986; Nolf and Cappetta, 1989). The Badenian otoliths (Borač: 2 specimen; Drnovice: 3 specimens) could be conspecific with the ones from the Mediterranean Lower Pliocene ( Nolf and Cappetta, 1989: pl. 3, figs 13-14). Both have a straight dorsal margin with rounded wavelike curves and a rather prominent anterodorsal angle. The same feature can be found in the fragmentary specimen of Nansenia sp. from the Mediterranean Tortonian ( Nolf and Steurbaut, 1983: pl. 1, fig. 13). This feature differentiates the Badenian otoliths from those of the Recent N. groenlandica (Reinhardt, 1840) (compare, e.g. Nolf and Steurbaut, 1983: pl. 1, fig. 12; Nolf, 2013: pl. 41), which is also reported from the Mediterranean Gelasian in Greece (and associated with a cold event; Agiadi et al., 2010, 2011) and the Pleistocene in Italy ( Girone, 2003; Girone et al., 2006) and the Indo-Pacific N. ardesiaca Jordan and Thompson, 1914 ( Ohe, 1985: p. 45, fig. 27). This feature, however, seems to vary in N. macrolepis (Gilchrist, 1922) – see Smale et al. (1995: pl. 9, fig. D1-2) and in the Nansenia sp. figured by Rivaton and Bourret (1999: pl. 88, figs 1-4). More and well-preserved fossil otoliths and data on Recent species are required to make more precise taxonomic conclusions.

The genus Nansenia includes foremost oceanic bathyph- ile pelagic fishes. Nansenia iberica Matallano, 1985 and N. oblita (Facciola, 1887) , e.g. both also living in the Mediterranean, are mesopelagic. At present, N. groenlandica lives in deep waters (up to 1400 m) in the North Atlantic and the Western Indian Ocean ( Froese and Pauly, 2016). The present fossil record is the first one of the genus in the Paratethys.

Sagamichthys ? sp. ( Pl. 1 , Fig. 6a, b). An otolith of a juvenile fish from Voděrady (L = 1.8 mm; H = 1.2 mm; L:H = 1.5) represents an alepocephalid fish. Its general morphology and sulcus show similarities with those of the present-day species Persparsia kopua (Phillips, 1942) – see Smale et al. (1995: pl. 10, fig. G1) – and with Sagamichthys abei Parr, 1953 ( Nolf, 2013: pl. 42). The former species is a mesopelagic fish distributed everywhere in the southern hemisphere (excluding South America) in the depth range of 700 to 1500 m, whereas the latter lives in the Pacific. In comparison with the Eastern Atlantic (see Smale et al., 1995: pl. 10, figs H, I) and Tortonian ( Lin et al., 2015: pl. 2, fig. 4) S. schnakenbecki (Krefft, 1953) , the Badenian specimen is clearly oblique along a praeventral-postdorsal axis. Otoliths of the West Indian S. gracilis Sazov, 1978 , are unknown. The available fossil material does not allow any more precise systematic conclusions.

Figures

Plate 1. – Explanation of the plates:All figured specimens are deposited in the collections of the Department of Geological Sciences, Masa- ryk University (DGS MU), Brno, Kotlářská 2, Czech Republic.The fossil otoliths bear numbers of the collection of types and figured fossil fish specimens in the DGS MU (Inv. Nr. O 315-377). The annotations a, b and c are used to indicate ventral, inner (= mesial) and posterior views, respectively. Figures with only numbers and no letter show inner views. In the captions, L stands for left otolith and R for right oto- lith. 1: Conger aff. conger (Linnaeus, 1758); Drnovice, Badenian, L (DGS MU Inv. Nr. O 315). 2: Serrivomer sp.; Lomnička, Badenian, R (DGS MU Inv. Nr. O 316). 3: Pterothrissus umbonatus (Koken, 1884); Drnovice, Badenian, R (DGS MU Inv. Nr. O 317). 4: Bathycongrus nagymarosyi Nolf & Brzobohaty, 1994; Voděrady, Badenian, R (DGS MU Inv. Nr. O 318). 5: Glossanodon cf. leioglossus (Valenciennes, 1848); Lomnice u Tišnova, Badenian, R (DGS MU Inv. Nr. O 319). 6: Sagamichthys? sp.; Voděrady, Badenian, R (DGS MU Inv. Nr. O 320). 7: Sarmatella aff. pulchra (Smigielska, 1966); Lomnice u Tišnova, Badenian, L (DGS MU Inv. Nr. O 321). 8: Scopelarchus aff. guentheri Alcock, 1896; Hrušovany nad Jevišovkou, Badenian, R (DGS MU Inv. Nr. O 326). 9: Etrumeus? weileri (Smigielska, 1966); Borač, Badenian, R (DGS MU Inv. Nr. O 323). 10: Bregmaceros albyi (Sauvage, 1880); Lomnice u Tišnova, Badenian, R (DGS MU Inv. Nr. O 324). 11: Physiculus aff. huloti Poll, 1953; Lomnice u Tišnova, Badenian, L (DGS MU Inv. Nr. O 325). 12: Nansenia sp.; Borač, Badenian, L (DGS MU Inv. Nr. O 322). 13: Merluccius aff. merluccius (Linnaeus, 1758); Lomnice u Tišnova, Badenian, L (DGS MU Inv. Nr. O 327).