Cretopachyderes burmitinus, Kundrata & Prosvirov & Long & Packova, 2021

Kundrata, Robin, Prosvirov, Alexander S., Long, Roger & Packova, Gabriela, 2021, Cretopachyderes gen. nov., a new remarkable click beetle (Coleoptera: Elateridae: Agrypninae) from the mid-Cretaceous Burmese amber, Fossil Record 24 (2), pp. 347-355 : 349-351

publication ID

https://doi.org/ 10.5194/fr-24-347-2021

publication LSID

lsid:zoobank.org:pub:1894924B-A1FE-43AF-A97F-50DAE387E9C0

persistent identifier

https://treatment.plazi.org/id/428E20BE-1DC2-482F-B81C-A8BC1439A027

taxon LSID

lsid:zoobank.org:act:428E20BE-1DC2-482F-B81C-A8BC1439A027

treatment provided by

Felipe

scientific name

Cretopachyderes burmitinus
status

sp. nov.

Cretopachyderes burmitinus sp. nov.

urn:lsid:zoobank.org:act:428E20BE-1DC2-482F-B81C-A8BC1439A027 ; Figs. 1–4 View Figure 1 View Figure 2 View Figure 3 View Figure 4 .

Type material

Holotype, adult specimen, sex unknown, MAM0012 (MAM). A complete beetle is included in a transparent, yellow amber piece with dimensions of 13.5 × 15.9 × 4 mm, without any syninclusions, only with air bubbles.

Type stratum and age

Mid-Cretaceous (upper Albian to lower Cenomanian) ( Shi et al., 2012; Mao et al., 2018; Yu et al., 2019).

Type locality

Hukawng Valley, Tanai Township, Myitkyina District, Kachin State, northern Myanmar.

Etymology

The specific epithet “ burmitinus ” is derived from “burmite” (mid-Cretaceous Burmese amber).

Diagnosis

As for the genus (vide supra).

Description

Adult. Body ( Fig. 1 View Figure 1 ) about 7.5 mm long (measured from frons to apex of elytra) and 2.6 mm wide (measured across widest part of pronotum), oval-elongate, rather flattened; dorsally sparsely setose.

Head ( Figs. 1–3 View Figure 1 View Figure 2 View Figure 3 ) subquadrate, retracted to prothorax, 0.45 times as wide as pronotum width. Eyes moderately large, strongly protuberant, with their major part hidden in prothorax. Labrum wider than long, anteriorly slightly concave. Antenna ( Figs. 2b View Figure 2 ; 3c–e View Figure 3 ) serrate from antennomere 4, relatively short, almost reaching posterior angles of pronotum; scape robust; pedicel not well visible; antennomere 3 short, slightly longer than wide, antennomere 4 1.6 times as long as antennomere 3, antennomeres 4–10 elongate, with short serrations; terminal antennomere simple, elongate, about 3 times as long as wide, about 1.3 times as long as antennomere 10, widest at two-thirds, apically narrowed and narrowly

350 R. Kundrata et al.: Cretopachyderes gen. nov., a new remarkable click beetle rounded. Mandible robust, relatively short, not well visible. Maxillary palpus moderately long; terminal palpomere securiform, apically flattened.

Pronotum ( Figs. 1–3 View Figure 1 View Figure 2 View Figure 3 ) weakly convex, without posterior angles widest at one-third, 1.45 times wider than long if measured along midline, and when posterior angles included then widest between their apices, and about 1.1 times longer than wide if length measured between anterior and posterior angles and width between the apices of posterior angles. Anterior margin widely concave; anterior angles very short; lateral sides rounded anteriorly, then sub-parallel; posterior angles very long, about 0.7 times as long as pronotal length when measured along midline, surpassing humeral part of elytra, gradually narrowed toward apex, relatively sharp, oriented posteriorly, with inner margin with distinct tooth and outer margin slightly concave; posterior margin medially emarginate. Lateral carina distinct, complete. Disc rather flattened, distinctly punctate; punctures sparser medially and denser and larger on sides, interstices smooth; sublateral carina distinct, coming from subapical inner part of posterior angle and running medially across whole length of posterior angle. Hypomeron somewhat shiny, with posterior margin subapically subrectangularly emarginate, with punctures distinct, large, separated 1.0–2.0 times of their diameter. Pronotosternal sutures almost straight, anteriorly open. Prosternum ( Figs. 2b View Figure 2 ; 3c View Figure 3 ) elongate, about 2.7 times as long as wide including prosternal process, about 1.8 times as long as wide without prosternal process; prosternal lobe well developed, short, rounded; prosternal process robust, very slightly gradually narrowed toward apex, subapically abruptly narrowed. Scutellar shield ( Figs. 1b View Figure 1 ; 2a View Figure 2 ; 3a View Figure 3 ; 4a View Figure 4 ) about 1.5 times as long as wide, narrowed subanteriorly, widest medially, then gradually narrowed toward apex, widely rounded anteriorly, narrowly rounded posteriorly, sides sinuate. Elytra ( Figs. 1a, c View Figure 1 ; 2a, b View Figure 2 ; 3a, c View Figure 3 ; 4a, b View Figure 4 ) together about twice as long as wide (measured at widest place) and 2.5 times as long as pronotum; weakly convex, widest at basal third, then gradually slightly narrowed towards apex; each elytron basally distinctly but weakly emarginate, with striae formed by lines of fine punctures; interstriae smooth, slightly convex; pubescence sparse, semi-erect, moderately long, oriented apically; elytral epipleuron moderately narrowed behind metacoxae. Mesoventrite not well visible. Metaventrite rather short and wide, 1.25 times as wide as long, moderately convex, sparsely covered by punctures. Metacoxal plate ( Fig. 2b View Figure 2 ) with basal portion well developed, wide, posteriorly widely roundly emarginate, then obliquely abruptly narrowed towards body edge. Hind wing not visible. Leg ( Figs. 1a–c View Figure 1 ; 2a–b View Figure 2 ; 4d–h View Figure 4 ) slender, femur robust, elongate, ventrally with distinct groove for reception of tibia; tibia shorter than femur, densely covered with thick long sharp setae, without distinct spurs; tarsus slender, elongate, distinctly longer than tibia. Tarsomere I longest, slightly shorter than tarsomere II and III combined; length ratio of tarsomeres I–V 4.2: 2.7: 2.0: 1.0: 3.0; tarsomeres I–IV gradually shorter, simple, almost subparallel sided, only apically slightly widened, moderately densely covered by setae, which are denser and longer apically; apical tarsomere elongate, slender; claws ( Fig. 4h View Figure 4 ) simple, slender, long, moderately curved.

Abdomen ( Figs. 2b View Figure 2 ; 4c View Figure 4 ) with ventrites moderately densely covered by distinct punctures; ventrites 1–4 subequal in length; ventrite 5 about 1.35 times as long as ventrite 4, subtriangular, narrowly rounded to subacute apically. Pregenital segments and genitalia not visible.

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