Thyropygus implicatus ( Demange, 1961 ) Demange, 1961

Pimvichai, Piyatida, Enghoff, Henrik & Panha, Somsak, 2009, A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae), Zootaxa 2016, pp. 17-50 : 31-32

Thyropygus implicatus ( Demange, 1961) , n. comb.

( Figs. 2 , 3 , 8 A–D)

Cornugonus implicatus Demange, 1961: 181 . Holotype (not seen) in the Natural History Museum, London.

Material: 10 males, 5 females, and 1 juvenile THAILAND, Satun Province, Thale Ban National Park, 6° 42ˏ 29˝ N, 100° 10ˏ 46˝ E. 15 October 1991. M. Andersen, O. Martin and N. Scharff leg., ( ZMUC).

Diagnosis: A species of the opinatus subgroup. Spatulate lobe (sl) at the apical part of telopodite terminating in a sharp brown spine. Similar in this respect to T. opinatus , T. floweri and T. erectus . Differs from these species by the having the lateral process of the anterior coxal fold (alp) apically bent abruptly mesad, its tip with serrate margins.

Description: Adult males with 63–70 podous rings, no apodous rings. Length ca. 12–13 cm, width ca. 7.4–8.0 mm. Adult females with 65–80 podous rings, no apodous rings. Length ca. 12–22 cm, width ca. 8.0–11.0 mm. Color in life unknown; preserved specimens with head, antennae, legs, prozona, middorsal metazona, epiproct, paraprocts and hyproproct brown; metazona, margins of epiproct and paraprocts, and the tip of hypoproct darker brown.

Gonopods ( Figs. 8 A–D): Anterior coxal fold (ac) ( Fig. 8 A): lateral process (alp) apically bent abruptly mesad, tip with serrate margins, posterior side with a horizontal crest; mesal process (amp) much shorter than lateral process, directed meso-distad, pointed. Posterior coxal fold (pc) ( Fig. 8 B) with lateral paracoxites (px), digitiform; mesal process (pmp) complex, strongly developed along an anterior-posterior axis, with a large posterior broad lobe (pbl) and an anterior slender digitiform process (adp); lateral process (plp) small, pointed. Telopodite ( Figs. 8 C–D) leaving coxite between pmp and plp; femoral spine (fe) basally massive, ending in sharp point, directed almost straightly mesad, in situ resting against posterior surface of alp, fitting under its horizontal crest, crossing over with fe of the opposite side; telopodite distally to fe with a large, round lobe (lo) projecting distolaterally; tibial spine (ti) long, slender, curving in horizontal plane, its tip upturned and resting against anterior surface of adp; apical part: spatulate lobe (sl) terminating in a sharp brown spine; palette (pa) simple, gutter-like, distally with about nine brownish blepharochaetae (bp).

Distribution ( Fig. 20): Described from MALAYSIA, Penang Hill (5° 25ˏ 29˝ N, 100° 16ˏ 7˝ E) ( Demange, 1961), based on specimens (including holotype) belonging to the Natural History Museum in London, and further specimens from MALAYSIA, Malacca, Sultanate of Kedah (2° 27ˏ 16˝ N, 102° 25ˏ 47˝ E) belonging to Naturalis, Leiden. The species was recorded as new for the Thai fauna by Enghoff (2005) based on the specimens studied by us ( THAILAND, Satun Province, Thale Ban National Park).

Figures

FIGURE 2. Thyropygus implicatus, SEM. A: apical view, left telopodite removed. B: posterior view, left telopodite removed. C: lateral view, with right telopodite. D: left coxa, lateral view.

FIGURE 3. Thyropygus implicatus, SEM. Spatulate lobe (sl), palette (pa), blepharochaetae (bp).

FIGURE 8. Thyropygus implicatus, specimen from Thale Ban National Park, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view.