Calityla trinitaria, Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 20

Calityla trinitaria , new species

Figs. 58–66

Types: Male holotype and female paratype from CALIFORNIA: Trinity Co., Indian Valley , parking area for Indian Creek Valley Cave , elev. 700 m., 40.5634°N, - 123.4197°W, collected 27 October 1990, by D. Ubick, W. Rauscher ( CAS) .

Diagnosis: Taiyutyla trinitaria is sympatric, possibly syntopic, with the previous species but may easily be distinguished from it by the far more complicated coxites of the male posterior gonopods, and the femoral knobs of the male pregonopodal legs, which are on legpairs three to seven, while in T. ubicki the knobs are found only on legpairs six and seven.

Etymology: The species name is an adjective based on Trinity County.

Description: Male holotype: Length, 11.0 mm. 17 ocelli in triangular eyepatch. Metazonites with low, narrow shoulders on rings 4–26. Color pale cream-tan lightly mottled brown. Legpairs one and two reduced, three to seven enlarged, with moderate angular knobs basal on femora three and to seven ( Figs. 61–66 ). Anterior gonopods ( Fig. 58 ) simple, blunt, roughened apicolaterally with small cuticular teeth ( Fig. 59 ). Posterior gonopod coxites bearing large, complex fimbriate branch distal of midlength, distally acuminate, hooked ( Fig. 60 ). Legpair 10 coxae of normal size, with small glands, legpair 11 femora with long, thin, dorsally directed knobs.

Female 11.0 mm long, similar to male in nonsexual characters.

Distribution: Known only from type locality.

Note: This species is clearly quite different from its sympatric congener T. ubicki .

Figures

FIGS. 51–66. Calityla spp. Figs. 51–57. Calityla ubicki. Fig. 51. Anterior gonopods, posterior view. Fig. 52. Left posterior gonopod coxite, posterior view. Figs. 53–57. Male legs 3–7, respectively. Figs. 58–66. Calityla trinitaria. Fig. 58. Anterior gonopods, posterior view. Fig. 59. Tip of left anterior gonopod, posterior view. Fig. 60. Left posterior gonopod coxite, posterior view. Figs. 61–66. Male legs 3–7, respectively. Fig. 67. Femoral knob of male leg 7.