Calityla essigi, Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 20

Calityla essigi , new species

Figs. 48–50 , 67–74

Types: Male holotype and four male paratypes from CALIFORNIA: Santa Cruz Co., Ben Lomond , ca. 40.56°N, - 123.42°W (estimated coordinates taken from Highlands County Park in Ben Lomond ), collected 22 January 1955, by D. Burdick, M. Wasbauer ( EMEC) .

Diagnosis: This species resembles the preceding one, but is distinct in the more massive fimbriate branch of the posterior gonopod coxites and their more robust terminal process, with the subterminal tooth displaced basally.

Etymology: The species epithet honors the late Edward O. Essig, founder of the Essig Museum of Entomology at the University of California, Berkeley. Dr. Essig shared with WAS an interest in the horticulture of the genus Iris .

Description: Paratype male from Ben Lomond: Length, 11.0 mm. 22 ocelli in triangular eyepatch. Metazonites with prominent angular shoulders on rings 2–27. Color chestnut brown marked darker purplish brown. Legpairs one and two reduced, three to seven enlarged, with mesal knobs on femora of legpairs three to seven; hardly discernible on femora 3, gradually enlarged on more posterior femora, large, squarish and angular on femora seven ( Figs. 70–74 ). Anterior gonopods ( Figs. 48, 49 , 67) with prominent mesobasal, single seta on each side, simple, anteroposteriorly flattened, with sinuate posterior ridge, small serrations on lateral sides near tips ( Fig. 49 ), slightly hooked apically. Posterior gonopod coxites ( Figs. 50 , 68, 69 ) bearing large, complex fimbriate branch distal of midlength, terminating in large bifurcate process that bears a tooth near its base. Legpair 10 coxae of normal size, with small glands, legpair 11 femora with long, thin, dorsally directed knobs.

Distribution: Known only from the type locality. Ben Lomond is a town; the same name is applied to a nearby mountain, and both are close to Big Bend Redwoods State Park.

Figures

FIGS. 45–50. Calityla spp. Figs. 45–47. Calityla siskiyou. Fig. 45. Left anterior gonopod, posterior view. Fig. 46. Tip of left posterior gonopod coxite, posterior view. Fig. 47. Right posterior gonopod coxite, posterior view. Fig. 48–50. Calityla essigi. Fig. 48. Right anterior gonopod, posterior view. Fig. 49. Tip of right posterior gonopod, posterior view. Fig. 50. Tip of left posterior gonopod coxite, mesoposterior view.

FIGS. 68–84. Calityla spp. Figs. 68–74. Calityla essigi. Fig. 68. Left posterior gonopod coxite, posterior view. Fig. 69. Tip of left posterior gonopod coxite, posterior view. Figs. 70–74. Male legs 3–7, respectively. Figs. 75–85. Calityla humboldtensis. Fig. 75. Anterior gonopods, posterior view. Fig. 76. Posterior gonopods, posterior view. Fig. 77. Tip of left posterior gonopod coxite, posterior view. Figs. 78–82. Male legs 3–7, respectively. Fig. 83. Male leg 10. Fig. 84. Male leg 11. Fig. 85. Femoral knob of male leg 7.