Calityla ubicki, Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 17-18

Calityla ubicki , new species

Figs. 51–57

Types: Male holotype from CALIFORNIA: Trinity Co., Indian Valley Creek Cave , 4 air mi SSE of Hyampom, elev. ca. 565 m., 40.567°N, - 123.416°W, collected 27 October 1990, by D. Ubick, W. Rauscher ( CAS) .

Diagnosis: Unlike any other, the femoral knobs of the male pregonopodal legs are limited to legpairs six and seven, with the knobs on prefemora seven basal, large and fungiform, and on prefemora six mesal and as a small swelling.

Etymology: The species epithet recognizes the collector, Darrell Ubick of the California Academy of Sciences.

Description: Male holotype: Length, 10.0 mm. 14 poorly formed and pigmented ocelli in triangular eyepatch. Metazonites with low shoulders on rings 5–24. Unpigmented. Legpairs one and two reduced, three to seven enlarged, with elongate tarsi, large fungiform knobs basal on femora seven, mesodistal low swellings on femora 6, knobs of femora seven roughened by small teeth ( Figs. 53–57 ). Anterior gonopods ( Fig. 51 ) simple, slender, tapering, slightly hooked apically, with subapical posterior tooth. Posterior gonopod coxites ( Fig. 52 ) bearing short, curved fimbriate branch midlength, terminating in small, bifurcate process. Legpair 10 coxae of normal size, with small glands, legpair 11 femora with blunt, dorsally directed knobs.

Distribution: known only from the type locality.

Notes: C alityla ubicki would seem to have some adaptation to cave life, with reduced eyes and pigment and somewhat elongate leg tarsi.

Figures

FIGS. 51–66. Calityla spp. Figs. 51–57. Calityla ubicki. Fig. 51. Anterior gonopods, posterior view. Fig. 52. Left posterior gonopod coxite, posterior view. Figs. 53–57. Male legs 3–7, respectively. Figs. 58–66. Calityla trinitaria. Fig. 58. Anterior gonopods, posterior view. Fig. 59. Tip of left anterior gonopod, posterior view. Fig. 60. Left posterior gonopod coxite, posterior view. Figs. 61–66. Male legs 3–7, respectively. Fig. 67. Femoral knob of male leg 7.