Calityla siskiyou, Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 17

Calityla siskiyou , new species

Figs. 37–47

Types: Male holotype, male paratype and eight female paratypes from CALIFORNIA: Shasta Co., 8 mi S of Dunsmuir, Shasta-Trinity National Forest , estimated coordinates measured 8 mi S of Dunsmuir along I-5, 41.1095°N, - 122.3225°W, collected 23 November 1954, by E. Gilbert & R. Schuster ( EMEC); county recorded as Siskiyou County on the label, but the Shasta County line is only 2 mi S of Dunsmuir.

Diagnosis: This species seems quite close to T. ubicki , but differs in details of the posterior gonopod coxites and in having femoral knobs on pregonopodal legpairs three to seven, instead of just six and seven.

Etymology: The name is a noun in apposition, referring to the occurrence of the species in Siskiyou County.

Description: Male holotype: Length, 10.0 mm. 19 ocelli in triangular eyepatch. Metazonites with low, narrow shoulders on rings 6–24. Color uniform pale tan (after long preservation). Legpairs one and two reduced, three to seven enlarged, with moderate angular knobs basal on femora three and four, mesodistal on femora five, distal on femora six and seven ( Figs. 40–44 ). Anterior gonopods ( Figs. 37 , 45 ) with prominent mesobasal, single seta on each side, simple, slender, tapering, slightly hooked apically, with subapical posterior tooth that is slightly hamate. Posterior gonopods with relatively small coxites ( Figs. 38, 39 , 46, 47 ) bearing large, complex fimbriate branch (fb, Fig. 46 ) distal of midlength, terminating in elongate bifurcate process that bears a basal tooth. Legpair 10 coxae of normal size, with small glands, legpair 11 femora with long, thin, dorsally directed knobs.

Female 10 mm long, similar to male in nonsexual characters.

Distribution: Known only from the type locality.

Figures

FIGS. 30–44. Taiyutyla amicitia and Calityla siskiyou. Figs. 30–36. Taiyutyla amcitia. Fig. 30. Anterior gonopods, posterior view. Fig. 31. Left posterior gonopod coxite, posterior view. Figs. 32–36. Male legs 3–7, respectively. Figs. 37–44. Calityla siskiyou. Fig. 37.Anterior gonopods, posterior view. Fig. 38. Right posterior gonopod coxite, posterior view. Fig. 39. Tip of right posterior gonopod coxite, posterior view. Figs. 40–44. Male legs 3–7, respectively.

FIGS. 45–50. Calityla spp. Figs. 45–47. Calityla siskiyou. Fig. 45. Left anterior gonopod, posterior view. Fig. 46. Tip of left posterior gonopod coxite, posterior view. Fig. 47. Right posterior gonopod coxite, posterior view. Fig. 48–50. Calityla essigi. Fig. 48. Right anterior gonopod, posterior view. Fig. 49. Tip of right posterior gonopod, posterior view. Fig. 50. Tip of left posterior gonopod coxite, mesoposterior view.