Taiyutyla jonesi ( Chamberlin, 1951 )

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 9-11

Taiyutyla jonesi ( Chamberlin, 1951)

Figs. 1–11

Conotyla jonesi Chamberlin 1951:6 . Type locality: Oregon, Eugene , Lane Co. The whereabouts of the male holotype are unknown and it is presumed lost.

Taiyutyla jonesi, Hoffman 1961:270 .

Taiyutyla corvallis Chamberlin 1952:1 . Type locality: Oregon, Corvallis. Location of male holotype unknown, presumed lost. Hoffman, 1961:270; Shear 1971:87, 1976:6. New Synonymy.

Taiyutyla lewisi Shear, 1976:13 . Type locality: Lane Co., 5 mi N of Elmira at the junction of Demming and Territorial Roads, male holotype and male and female paratypes (AMNH). New Synonymy.

New records: UNITED STATES: OREGON: Benton Co., Clemens Park, Seely Creek Road, 0.3 mi from SR-34, North Fork of Alsea River, elev. 115 m., 44.4089°N, - 123.5675°W, 4 December 2005, W. Leonard, C. Richart, 3mm f (CAS); Multnomah Co., Gabbert Butte, Gresham, elev. 255 m., 45.4730°N, - 122.4356°W, 18 December 2017, P. Nosler, m f; West Bliss Butte near Clackamas County line, elev. 140 m, 45.4633°N, - 122.4622°W, 18 November 2017, P. Nosler, 2mm; Towle Butte, near Gresham, elev. 160 m., 45.4665°N, - 122.4540°W, 12 January 2018, P. Nosler, 2mm; same as previous but elev. 260 m., 45.4665°N, - 122.4500°W, 2mm; same as previous but elev. 155 m., 45.4664°N, - 122.454106°W, 2mm; Linn Co.: Cartney Park Drive near McCartney Park and Boat Ramp, Harrisburg, elev. 90 m., 44.3169°N, - 123.2138°W, 23 November 2017, P. Nosler, m; Tillamook Co., Rogers Forest Park, Beaver Dam Road 0.4 mi W of SR-6, Gales Creek, Tillamook State Forest, elev. 495 m., 45.6150°N, - 123.3780°W, 6 January 2018, P. Nosler, m. WASHINGTON: Whatcom Co., White Cap Road at Chuckanut Drive, elev. 60 m., 48.6623°N, - 122.4922°W, 20 February 2004, C. Richart, from the litter and woody debris of a mixed forest including Acer macrophyllum , Alnus rubra , Thuja plicata , Tsuga heterophylla , Pseudotsuga menziesii , Polystichum munitum , 3mm 2ff (CAS); Pierce Co.: Sequalitchew Creek Trailhead, elev. 61– 67 m., 47.107°N, - 122.645°W, 28 January 2012, R.; Crawford, 2 mm 3 ff (UWBM). CANADA: BRITISH COLUMBIA: Vancouver Island, Nanaimo River Road, 4.1 mi W of BC-1, elev. 130 m., 49.0695°N, - 123.9547°W, 2 March 1983, B. D. Ainscough, 1m (CAS); Lone Tree Hill, Saanich, elev. 240 m., 48.5182°N, - 123.5118°W, 21 October 2004, K. Ovaska, m (CAS); Colwood, 48.4391°N, - 123.4504°W, undated, K. Ovaska, L. Sopuck, m f (CAS); Mattheson Lake Regional (Provincial) Park, Metchosin, elev. 55 m., 48.3604°N, - 123.5919°W, 22 October 2009, K. Ovaska, 2ff; Florencia Bay, Pacific Rim National Park, elev. 20 m., 49.0128°N, - 125.6522°W, 29 November 2003, K. Ovaska, L. Sopuck, 5mm (CAS); Goldmine, Pacific Rim National Park, 49.0115°N, - 125.6215°W, 27 November 2003, K. Ovaska, L. Sopuck, 2mm (CAS).

Notes: Direct comparison of specimens from numerous localities establishes that T. jonesi is the senior synonym and both T. corvallis and lewisi are junior to that name. The distribution of the species is unusual in that only two records are known from Washington, between the localities in Oregon and British Columbia. However, scanning electron micrographs reveal the surprisingly exact identity of the gonopods of specimens from both areas. The gonopods were illustrated by Shear (1971) as were those of T. corvallis and T. lewisi by Shear (1976). Examination of types and new specimens reveal that both these names are synonyms of T. jonesi . Further illustrations of the gonopods and pregonopodal legs of males are presented here ( Figs. 1–11 ).

Figures

FIGS. 1–4. Taiyutyla jonesi. Fig. 1. Anterior gonopods, posterior view. Fig. 2. Posterior gonopod coxites, posterior (right) and lateral (left) views. Fig. 3. Posterior gonopod coxites, posterior view. Fig. 4. Left posterior gonopod coxite, posterior view.

FIGS. 5–21. Taiyutyla spp. Figs. 5–11. Taiyutyla jonesi. Fig. 5. Anterior gonopods, posterior view. Fig. 6. Right posterior gonopod coxite, posterior view. Figs. 7–11. Male legs 3–7, respectively. Figs. 12–18. Taiyutyla Tillamook. Fig. 12. Anterior gonopods, posterior view. Fig. 13. Right posterior gonopod coxite, posterior view. Figs. 14–18. Male legs 3–7, respectively. FIGS. 19–21. Taiyutyla acuphora. Fig. 19. Left anterior gonopod, posterior view. Fig. 20. Left anterior gonopod tip, mesal view. Fig. 21. Right posterior gonopod coxite, posterior view.