Complicatella complicata ( Shear, 1974 ) Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 48-49

Complicatella complicata ( Shear, 1974) , new combination

Bollmanella complicata Shear 1974, p. 146 .

Figs. 192–193

New Records: WASHINGTON: Thurston Co., Black Lake and Belmore Road at 66 th Avenue, Tumwater, elev. 60 m., 46.9889°N, - 122.9659°W, 19 November 2003, W. Leonard, mm; Hospital Creek , above confluence with Skookumchuck River , 5 horiz mi S, 3 horiz mi E of Vail, elev. 245 m., 46.7732°N, - 122.5855°W, 15 December 2003, W. Leonard, K. McAllister, mm ff; Rainier Road SE, 3.0 mi NW of Military Road SE, Fort Lewis Military Reservation , elev. ca. 100 m., ca. 46.9351°N, - 122.7600°W, 29 November 2003, W. Leonard, m; The Evergreen State College campus, Beach Trail N of Driftwood Road , Olympia , elev. 30 m., 47.0791°N, - 122.9776°W, 26 January 2003, W. Leonard, mm ff; same as previous but 16 March 2003, mm; same as previous but 23 March 2003; same as previous but 22 December 2003, mm ff; Priest Point Park, Olympia, elev. ca. 65 m., ca. 47.07°N, - 122.89°W, 5 January 2003, W. Leonard, mm f; same as previous but 2 February 2003, m; same as previous but 14 November 2004, mm ff; along Rock Candy Mtn. Road, Capitol State Forest , elev. ca. 300 m., 47.0256°N, - 123.0735°W, 18 December 2004, W. Leonard, m. Grays Harbor Co. , Porter Creek Campground, Capitol State Forest , elev. 90 m., 46.9779°N, - 123.2565°W, 24 January 2005, W. Leonard, m. Cowlitz Co. , Germany Creek Road 5.5 mi N of Stella at SR-4, elev. 110 m., 46.2593°N, - 123.1344°W, 22 November 2003, W. Leonard, C. Richart, mm; same as previous but 8 December 2003, m f; same as previous but 11 November 2004, mm ff. Jefferson Co. , along Dosewallips River , Dosewallips Road 7.0 mi W of US-101, Olympic National Forest , elev. 115 m., 47.7326°N, - 123.0201°W, 19 November 2003, W. Leonard, m; Falls View Campground, US-101 7.8 mi NW of Brinnon, elev. 150 m., 47.7893°N, - 122.9255°W, 22 February 2003, W. Leonard, m f. Mason Co. , SR-119 6 mi W of US-101 at Hoodsport, elev. 310 m., 47.7326°N, - 123.2097°W, 7 February 2003, W. Leonard, mm. King Co. , FR-9034 near Granite Mountain Trailhead, Mt. Baker-Snoqualmie National Forest , elev. 580 m., 47.3977°N, - 121.4855°W, W. Leonard, 27 October 2007, mm f; same as previous but 26 March 2004, W. Leonard, C. Richart, m; same as previous but 25 October 2003. Lewis Co. , 604 Roswell Road, Centralia, elev. 65 m., 46.7227°N, - 122.9452°W, 25 January 2004, C. Richart, W. Leonard, mm ff; SE end of Riffle Lake , elev. 250 m., 46.4644°N, - 122.1705°W, 9 April 2004, W. Leonard, C. Richart, m; Slide Creek , Weyerhaeuser 4000 Rd 0.8 mi SW of Pe-Ell-McDonald Rd., elev. 190 m., 46.5250°N, - 123.1913°W, 4 December 2004, W. Leonard, C. Richart, mm ff; same as previous but 3 January 2005, W. Leonard; Stillman Basin, 1.8 mi on 4000 Weyerhaeuser from Pe-Ell McDonald Rd., elev. 290 m., 46.5145°N, - 123.2015°W, 4 December 2004, W. Leonard, C. Richart, from the litter and woody debris of a mixed forest including Thuja plicata , Alnus rubra , Rubus spectabilus , Pseudotsuga menziesii , and Polystichum munitum , m f; North Fork of Newaukum River , below confluence with Middle Fork, elev. 88 m., 46.6046°N, - 122.8481°W, 29 December 2004, W. Leonard, m. Notes : This species was described from 1 mi W of Bayshore and 4 mi N of Shelton, Mason Co. , Washington , and heretofore was known only from the type locality. New collecting revealed a much wider distribution, detailed above and in the supplemental KML map, and show this species to be the most widely distributed and most commonly collected conotylid in western Washington. All these populations are linked by identical anterior gonopods ( Fig. 192 ), but the posterior gonopod coxites (i.e., Fig. 193 ) are so complicated that it is difficult to view them from exactly the same angle and make comparisons. Thus it is not possible really to be sure that all collections belong to the same species. This can be worked out later with genetic data.

Descriptions of new species of Complicatella

Figures

FIGS. 190–193. Brunsonia chelanomagna and Complicatella complicata. Figs. 190, 191. Brunsonia chelanomagna. Fig. 190. Anterior gonopods, posterior view. Fig. 191. Left posterior gonopod coxite, posterior view. Figs. 192–193. Complicatella complicata. Fig. 192. Left anterior gonopod, posterior view. Fig. 193. Posterior gonopod coxites, posterior view.