Tuberculobasis cardinalis ( Fraser, 1946 ) Fraser, 1946

Machado, Angelo B. M., 2009, Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)., Zootaxa 2108, pp. 1-36 : 17

Tuberculobasis cardinalis ( Fraser, 1946) comb. nov.

Figures 47 , 95

Oxyagrion cardinalis Fraser (1946: 41–42); Soukup (1954: 12); St. Quentin (1960: 59); Kimmins (1966: 185). Oxyagrion cardinale: Rácenis (1959: 478) ; Costa (1978: 62); De Marmels (1984a: 24); Davies & Tobin (1984: 75); Bridges (1994: 44, 52); Tsuda (2000: 41); Costa et al. (2005: 23).

Leptobasis cardinalis: Von Ellenrieder & Garrison (2007: 25) .

Species examined. None.

Distribution. PERU, Loreto Department.

Remarks. T. cardinalis was described by Fraser (1946) under Oxyagrion based on one female specimen from Peru. Costa (1978) suggested that the species might belong in Acanthagrion due to the presence of thoracic stripes and De Marmels (1984a) provided a partial redescription of the holotype, including drawings and photographs. Von Ellenrieder & Garrison (2007) examined the holotype in BMNH and concluded that the species is not an Oxyagrion and should be assigned to Leptobasis . They provided drawings of its main diagnostic characters, some of which are reproduced here ( Figs 47 , 95 ). The presence of mound-like mesepisternal tubercles and ovipositor not surpassing tip of cercus ( Fig. 95 ), plus wing petiolation subequal to Ac justifies its transfer to Tuberculobasis . Von Ellenrieder & Garrison (2007) found some discrepancies between Fraser's description and the holotype that they regarded as unintentional mistakes by Fraser (1946). The most significant discrepancy was the presence of a vulvar spine mentioned by Fraser (1946) but missing in the holotype.

Male. Unknown.

Figures

FIGURES 47 – 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels].

FIGURES 95 – 100. Female S 8 – 10 in lateral view: T. cardinalis, holotype (95); T. mammilaris, Venezuela (96); T. costalimai, allotype (97); T. williamsoni, paratype (98); T. yanomami, paratype (99); Leptobasis vacillans, Venezuela (100) [95 by N. von Ellenrieder, 96, 99 by J. De Marmels].