Maietta sinistra, Santos & Santos & Ribeiro, 2019

Santos, Daubian, Santos, Rodrigo Dos Reis & Ribeiro, Guilherme Cunha, 2019, Systematics of the genus Maietta Alexander with description of three new species (Diptera: Tipulomorpha: Limoniidae), Zootaxa 4695 (4), pp. 331-350 : 341-345

Maietta sinistra sp. n.

( Figures 5e , 6e , 7e , 8e , 9e , 10e ).

Material examined. Holotype. Pinned specimen, male. Chile, Arauco, Nahuelbuta, Butamalal [37° 49’ S, 73° 14’ W], 1100–1400 m, 23/ 31-I-1954, Peña ( USNM). [The specimen designated here as the holotype of this new species was misidentified by Alexander as Maietta trimedia Alexander ] . Material identified by Charles Paul Alexan- der. Pinned specimens: Maietta trimedia Al. Det. C. P. Alexander , 1 female and 4 [sex unknown], Chile, Arauco, Nahuelbuta, Butamalal [37° 49’ S, 73° 14’ W], 1100–1400 m, 23/ 31-I-1954, Peña ( USNM) ; Maietta squamigera Al. Det. C. P. Alexander , 1 female, Chile, Arauco, Nahuelbuta, Pichinahuel [37° 48’ S, 73° 02’ W], 1100–1400 m, 23/ 31-I-1954, Peña ( USNM) ; Maietta trimedia Al. Det. C. P. Alexander , 1 [sex unknown], Chile, Arauco, Nahuelbuta, Pichinahuel [37° 48’ S, 73° 02’ W], 1200 m, 14/ 18-II-1956, Luis Peña ( USNM) . Specimens in slides prepared by Charles Paul Alexander: Maietta trimedia Al. Det. C. P. Alexander , male, Chile, Arauco, Nahuelbuta, Butamalal [37° 49’ S, 73° 14’ W], 1100–1400 m, 23/ 31-I-1954, Peña ( USNM) ; Maietta trimedia Al. Det. C. P. Alexander , female, Chile, Chiloé Is., Aucar [42° 10’ S, 73° 30’ W], 6/ 15-I-1952, Peña ( USNM) .

Etymology. The word “sinistra” is Latin and means “left”. This word was chosen in reference to the fact that both parameres are pointing to the left.

Diagnosis. Cercus smaller than one fifth of tergite X. Tergite X characteristic, being as wide as sternite IX. Both parameres pointed to the left.

Description. Head. Scape more cylindrical, 3.8 times longer than wide. Pedicel with circular shape, as long as wide. Scape 2.3 times longer than pedicel. Pedicel 1.5 times longer than first flagellomere. Penultimate flagellomere 1.8 longer than last one ( Fig. 5e ).

Thorax. Thoracic sclerites shown in Fig. 6e . Wing shown in Fig. 7e . Vein Sc ending near to R 2+3 but more distal, condition homoplastic with M. dextra sp. n. Vein R 2 weakly angulated and approximately 2.8 times longer than vein R 2+3. Length of Vein R 2+3+4 similar to vein R 2+3. First section of vein R 5 1.4 times longer than cross-vein r-m. First section of vein M 1+2 equal than cross-vein r-m. Three medial veins present. Vein M 3+4 equal in length to M 4, both 2.8 times the length of cross-vein m-cu. Distance between M 1+2 and R 5 equal to distance between R 4 and R 5 and to distance between M 3 and M 1+2.

Female terminalia. Female tergite X 1.5 times longer than wide. Dorsal margin of tergite X without projections. Cercus small, 4.8 times shorter than length of tergite X. Ventral margin of tergite IX 1.7 times longer than the dorsal margin. Dorsal projection of sternite IX somewhat angulated. Hypogynial valves almost circular, as long as wide, and 1.2 times wider than sternite IX. Posterior projection of sternite IX more stretched out ( Fig. 8e ).

Male Terminalia. Male terminalia illustrated in Fig. 9e (Details of chaetotaxy shown in Fig. 10e ). Gonocoxite 1.6 times longer than lobe and clasper of gonostylus, which are similar in length. Parameres with concave lateral margin. Sheath of aedeagus with bifid projection. The tips are 1.1 times smaller than the projection. Bridge between interbases wider than aedeagus. Long tip of bifid posterior projection of interbases 1.3 times longer than the short tip. Both projections of interbase pointing to the left.

Coloration. General coloration brown. Head: dark brown. Rostrum dark brown, palpi dark brown. Scape and pedicel brownish yellow, flagellum obscure yellow with outer segments lighter. Thorax: blackened brown, pleuron dark brown. Halters pale. Coxae and trochanters fulvous yellow, remainder of legs pale. Wings hyaline, scales light brown, veins nearly hyaline.Abdomen dark fulvous yellow with terminal segments dark brown but with hypogynial valves contrastingly yellow.

Remarks. Some specimens of this new species were misidentified by C. P. Alexander as M. trimedia Alexander and others as M. squamigera Alexander.

Figures

FIGURE 5. Head of Maietta (frontal view). All specimens male. a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: ant, antenna; ped, pedicel; plp, palpus; scp, scape.

FIGURE 6. Thorax of Maietta (lateral view). All specimens male. a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cerv scl, cervical sclerite; cx, coxa; hlt, halter; kepm, katepimeron; kepst, katepisternum; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtn, metanotum; p bas, posterior basalare; patg, paratergite; pprn, pospronotum; presct, prescutum; s scl, subalar sclerite; sct, scutum; sctl, scutellum.

FIGURE 7. Wing of Maietta (lateral view). All specimens male. a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n.

FIGURE 8. Female terminalia of Maietta (lateral view). a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexan- der, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: dp, dorsal projection; cer, cercus; hyp val, hypogynial valve; pp, posterior projection; st, sternite; tg, tergite.

FIGURE 9. Line drawings of the male terminalia of Maietta (dorsal view). a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: aed, aedeagus; c gonst, clasper of gonostylus; goncx, gonocoxite; int, interbase; l gonst, lobe of gonostylus; prm, paramere; sh aed, sheath of aedeagus.

FIGURE 10. Photographs of the male terminalia of Maietta (dorsal view). a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n.