Maietta dextra, Santos & Santos & Ribeiro, 2019

Santos, Daubian, Santos, Rodrigo Dos Reis & Ribeiro, Guilherme Cunha, 2019, Systematics of the genus Maietta Alexander with description of three new species (Diptera: Tipulomorpha: Limoniidae), Zootaxa 4695 (4), pp. 331-350 : 337-338

Maietta dextra sp. n.

( Figs 5c , 6c , 7c , 8c , 9c , 10c )

Material examined. Holotype. Pinned specimen, male. Chile, Arauco, Pillim Pilli [37° 46′ S, 73° 04′ W], 5– 800 m, 15-I-1954, Peña ( USNM). ( The specimen designated here as the holotype of this new species was misidentified by Charles Paul Alexander as Maietta trimedia Alexander ). Material identified by Charles Paul Alexander . Pinned specimens: Maietta squamigera Al. Det. C. P. Alexander , female, Chile, Arauco, Pillim Pilli [37° 46′ S, 73° 04′ W], 5– 800 m, 15-I-1954, Peña ( USNM) ; Maietta squamigera Al. Det. C. P. Alexander , male, Chile, Arauco, Nahuelbuta , Butamalal [37° 49’ S, 73° 14’ W], 1400 m, 23/ 31-I-1954, Peña ( USNM) . Specimens in slides prepared by Charles Paul Alexander : Maietta squamigera Al. Det. C. P. Alexander , female, Chile, Arauco, Nahuelbuta , Caramavida [37° 40’ S, 73° 20’ W], 1000 m, 5/ 10-II-1953, Peña ( USNM) ; Maietta trimedia Al. Det. C. P. Alexander , male, Chile, Arauco, Nahuelbuta , Butamalal [37° 49’ S, 73° 14’ W], 1100– 1400 m, 23/ 31-I-1954, Peña ( USNM).

Etymology. The word “dextra” is Latin and means “right”. This word was chosen with reference to the fact that both parameres are pointing to the right.

Diagnosis. Vein Sc ending distal to the end of CuA. Vein M 4 longer than vein M 3+4. Sheath of aedeagus with trifid tip. Lateral margin of paramere not bifurcated.

Description. Head. Scape piriform, 2 times longer than wide. Pedicel almost circular, as long as wide. Scape 1.5 times longer than pedicel. Pedicel 1.5 times longer than first flagellomere. Penultimate flagellomere 1.6 longer than last one ( Fig. 5c ).

Thorax. Thoracic sclerites shown in Fig. 6c . Wing shown in Fig. 7c . Vein Sc ending distal to R 2+3 and at level of end of CuA. Vein R 2 approximately 6 times longer than vein R 2+3. Vein R 2+3+4 3.6 times longer than vein R 2+3. Crossvein r-m 1.8 longer than first section of vein R 5. First section of vein M 1+2 1.2 times longer than cross-vein r-m. Three medial veins present. Vein M 3+4 shorter than M 4, similar in length to cross-vein m-cu. Distance between M 1+2 and R 5 similar to distance between R 4 and R 5, and greater than distance between M 3 and M 1+2.

Female terminalia. Female tergite X 2 times longer than wide. Dorsal margin tergite X with a projection on the anterior section near to membranous portion of the cercus. Cercus straight, almost half the length of tergite X. Dorsal projection of sternite IX angulated. Hypogynial valves almost circular, as long as wide, and less wide than sternite IX. Posterior projection of sternite IX zigzagged ( Fig. 8c ).

Male terminalia. Male terminalia illustrated in Fig. 9c (Details of chaetotaxy shown in Fig 10c ). Gonocoxite 2.3 times longer than lobe of gonostylus and 1.7 times longer than clasper of gonostylus. Clasper of gonostylus longer than lobe. Parameres with pointed anterior margin. Sheath of aedeagus with trifid projection. Size of tips 3.15 times smaller than the size of projection. Bridge between interbases with a width similar to that of aedeagus. Long tip of bifid posterior projection of interbases almost twice as long as the short tip. Both projections of dorsal interbase pointing to the right in dorsal view.

Coloration. General coloration dark brown. Head: brown. Rostrum dark brown, palpi brown. Scape and pedicel dark brown, flagellum obscure yellow with outer segments darker. Thorax: brown, pleuron testaceous. Halters pale with knobs light ochreous. Coxae and trochanters light brown, remainder of legs yellowish brown. Wings hyaline, scales light brown, veins nearly hyaline. Abdomen brown.

Remarks. Some specimens of this new species were misidentified by C. P. Alexander as M. trimedia Alexan- der, and as M. squamigera Alexander.

Figures

FIGURE 5. Head of Maietta (frontal view). All specimens male. a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: ant, antenna; ped, pedicel; plp, palpus; scp, scape.

FIGURE 6. Thorax of Maietta (lateral view). All specimens male. a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cerv scl, cervical sclerite; cx, coxa; hlt, halter; kepm, katepimeron; kepst, katepisternum; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtn, metanotum; p bas, posterior basalare; patg, paratergite; pprn, pospronotum; presct, prescutum; s scl, subalar sclerite; sct, scutum; sctl, scutellum.

FIGURE 7. Wing of Maietta (lateral view). All specimens male. a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n.

FIGURE 8. Female terminalia of Maietta (lateral view). a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexan- der, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: dp, dorsal projection; cer, cercus; hyp val, hypogynial valve; pp, posterior projection; st, sternite; tg, tergite.

FIGURE 9. Line drawings of the male terminalia of Maietta (dorsal view). a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n. Abbreviations: aed, aedeagus; c gonst, clasper of gonostylus; goncx, gonocoxite; int, interbase; l gonst, lobe of gonostylus; prm, paramere; sh aed, sheath of aedeagus.

FIGURE 10. Photographs of the male terminalia of Maietta (dorsal view). a: Maietta squamigera Alexander, 1929; b: Maietta trimedia Alexander, 1967; c: Maietta dextra sp. n.; d: Maietta edwardsi sp. n.; e: Maietta sinistra sp. n.